Rosulabryum bornholmense

(Winkelmann & R. Ruthe) J. R. Spence

Novon 19: 398. 2009.

Basionym: Bryum bornholmense Winkelmann & R. Ruthe Hedwigia 38(Beibl.): 120. 1899
Treatment appears in FNA Volume 28. Treatment on page 180. Mentioned on page 179.

Plants small, red-green. Stems 0.5–2 cm, fertile stems evenly foliate to weakly rosulate, innovations evenly foliate. Leaves of main rosette and innovations similar, innovation leaves smaller; somewhat irregularly twisted to contorted when dry, erect-spreading when moist, ovate, weakly concave, 1–2 mm; base not decurrent; margins recurved to mid leaf, weakly but distinctly serrulate distally, limbidium moderately distinct, of 1 or 2 rows of cells; apex acute; costa excurrent, awn pigmented; proximal laminal cells short-rectangular; medial and distal cells rhomboidal, 14–20 µm wide, 3–4: 1, walls thick, not porose. Specialized asexual reproduction by rhizoidal tubers, amber, orange, orange-red, or dark red, becoming orangebrown, pale-brown, or dark-brown when old, usually brighter than rhizoids, (120–) 180–350 µm, cells not or only weakly protuberant. Sexual condition dioicous, synoicous, or autoicous. Capsule nutant, red to redbrown, cylindric, 2–3 mm.


Phenology: Capsules mature Apr–Aug.
Habitat: Disturbed soil
Elevation: low elevations (100 m)

Distribution

V28 287-distribution-map.gif

Introduced; Calif., Europe

Discussion

A. C. Crundwell and H. L. K. Whitehouse (2001) revised Rosulabryum bornholmense, providing new criteria to separate it from the closely related R. rubens. In addition to tuber and awn differences, the cells of the tubers in R. bornholmense are 45–60 µm wide, while those of R. rubens are 30–35 µm wide. Most North American collections can be referred to the latter species.

Selected References

None.

Lower Taxa

None.

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... more about "Rosulabryum bornholmense"
dark-brown +, pale-brown +, orangebrown +, dark red +, orange-red +  and orange +
usually brighter +
entire to distinctly +
John R. Spence +
(Winkelmann & R. Ruthe) J. R. Spence +
pigmented +
reduced +
curved +  and straight +
not decurrent +
Bryum bornholmense +
cucullate +
red +  and redbrown +
long-exserted +
cylindric +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
protuberant +
autoicous +, synoicous +, dioicous +  and sexual +
quadrate +  and long-rectangular +
not differentiated +
appendiculate +
subpercurrent +
hexagonal;rhomboidal +
Calif. +  and Europe +
low elevations (100 m) +
hyaline;pale-yellow +
not adherent +
Disturbed soil +
not differentiated +
concave +  and ovate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
polyoicous +, monoicous +, dioicous +  and sexual +
imbricate;variously contorted +
erect;erect-spreading +
spathulate +, obovate +, ovatelanceolate +, ovate +  and lanceolate +
vegetative +
1 (?) +, 3 (?) +  and 4 (?) +
short-rectangular +  and quadrate +
short-conic +  and umbonate +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature Apr–Aug. +
differentiated +
smooth;papillose +
straight +
elongate +
dark yellow +
germinating +
erect +  and erect-spreading +
concave +, flat +, spathulate +, obovate +  and ovate +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Bryum sect. Trichophora +
Rosulabryum bornholmense +
Rosulabryum +
species +
lanceolate;narrowly lanceolate +
distinct +
straight +
specialized +
autoicous +, polyoicous +, synoicous +, dioicous +  and sexual +
sinuate +