Quercus garryana var. breweri

(Engelmann) Jepson

Fl. Calif. 1(2): 354. 1909.

EndemicIllustrated
Basionym: Quercus breweri Engelmann in S. Watson, Bot. Calif. 2: 96. 1880
Treatment appears in FNA Volume 3.

Shrubs, spreading and clonal, to 2-3 m, multitrunked. Twigs reddish, sparsely puberulent, often glabrate, without spreading hairs. Buds reddish-brown, ovoid, 2-5 mm, glandular-puberulent. Leaf-blade abaxially light green, velvety to touch, sparsely to densely covered with erect (2-) 4-6-rayed hairs 0.2-0.5 mm.


Phenology: Flowering late spring–early summer.
Habitat: Montane conifer forests and chaparral
Elevation: 1400-1900 m

Discussion

Quercus garryana var. breweri appears to be endemic to the Siskiyou region of California and Oregon; it may extend into the northern Sierra Nevada of California. Specimens sometimes placed here from the Coast Ranges of northern California are probably shrubby forms of Q. garryana var. garryana or hybrids between the latter and Q. durata (see treatment). It should be noted that key characteristics separating Q. garryana vars. breweri and semota from var. garryana (clonal habit, smaller, glabrate, brown buds, montane habitat of the former two varieties) suggest a relationship of these two varieties with the Rocky Mountain Q. gambelii. The latter species has smaller fruit than Q. garryana, but the extent of variation in this characteristic in Q. garryana var. breweri is unknown.

Selected References

None.

Lower Taxa

None.
puberulent +, glabrous +, pubescent +  and glandular-puberulent +
rounded +, blunt +  and acute +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Kevin C. Nixon +
(Engelmann) Jepson +
white;light gray +
subcordate;cuneate;rounded-attenuate +
unequal +
Quercus breweri +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
reddish-brown +
glandular-puberulent +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
2-6-lobed +
pistillate +, capitate +  and spicate +
starchy +  and fleshy +
hemispheric +, cup-shaped +  and saucer--shaped +
×12-22 +, 4mm +  and 10mm +
multibracteate +, spiny +  and scaly +
Calif. +  and Oreg. +
1400-1900 m +
pistillate +  and staminate +
biennial +, annual +  and maturation +
1-seeded +  and winged +
in groups +  and enclosed +
Montane conifer forests and chaparral +
erect +  and spreading +
pistillate +, spicate +  and staminate +
tomentulose +  and glabrate +
12 cm120 mm <br />0.12 m <br /> (14 cm140 mm <br />0.14 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (12 cm120 mm <br />0.12 m <br />) +
subrotund +  and moderately deeply lobed +
arranged +  and alternate +
blunt +, rounded +, obtuse +, spatulate +  and oblong +
entire +, dentate +  and serrate +
toothed +  and entire +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
light-brown +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
oblong +  and globose +
1.4 cm14 mm <br />0.014 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Flowering late spring–early summer. +
reddish-brown +  and yellowish +
connate +  and distinct +
tomentulose +  and canescent +
few-to-many +
tuberculate +
long-acute +
reduced +
anastomosing +, branching +  and unbranched +
dark green +  and yellowish +
glossy +  and bright +
Endemic +  and Illustrated +
inconspicuous +
distinct +
subulate +  and dilated +
enlarged +
yellowish +, waxy +  and light green +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Quercus douglasii var. neaei +, Quercus garryana var. jacobi +, Quercus jacobi +, Quercus lobata var. breweri +  and Quercus neaei +
Quercus garryana var. breweri +
Quercus garryana +
variety +
spheric +  and ovoid terete or angled +
bristle-tipped +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
glabrate +  and puberulent +
1,500 cm15,000 mm <br />15 m <br /> (2,000 cm20,000 mm <br />20 m <br />) +
shrub +  and clonal +