Ptychostomum pseudotriquetrum

(Hedwig) J. R. Spence & H. P. Ramsay ex Holyoak & N. Pedersen

J. Bryol. 29: 120. 2007.

Illustrated
Basionym: Mnium pseudotriquetrum Hedwig Sp. Musc. Frond., 190. 1801 Bryum pseudotriquetrum (Hedwig) P. Gaertner B. Meyer & Scherbius
Treatment appears in FNA Volume 28. Treatment on page 164. Mentioned on page 156, 158, 162.

Plants in dense turfs, green, red-green, or yellow-green, older shoots becoming redbrown. Stems 2–4 (–6) cm, tufted, weakly comose to evenly foliate, innovations elongate and evenly foliate; usually strongly radiculose well toward stem apices. Leaves green, red-green, or yellow-green, dull brown-red or brick-red with age, somewhat twisted to contorted when dry, ovate, flat to weakly concave, (1–) 2–3 (–4) mm, not much enlarged toward stem apex; base strongly and narrowly decurrent; margins revolute to mid leaf or beyond, limbidium strong, in 2 or 3 rows; apex acute; costa short-excurrent, awn smooth; proximal laminal cells short-rectangular, 3: 1, same width or somewhat wider than more distal cells; medial and distal cells rhomboidal, 14–22 µm wide, (2–) 3: 1, walls firm to often incrassate. Specialized asexual reproduction occasional, by leaf-axil gemmae, brown. Sexual condition dioicous. Seta 1–3 (–4) cm. Capsule brown, elongate-ovate, symmetric, 3–5 mm, mouth yellow; operculum conic, apiculate; peristome well developed; exostome teeth yellow basally, hyaline distally, lamellae usually straight mid-tooth, pores absent along mid line; endostome not adherent to exostome, basal membrane high, 1/2 exostome height, segments with ovate perforations, cilia long, appendiculate. Spores 12–18 µm, finely papillose, pale-yellow or green.


Phenology: Capsules mature Apr–Oct.
Habitat: Wet soil, soil over rock, rock, fens
Elevation: low to high elevations (0-4000 m)

Distribution

V28 261-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., P.E.I., Que., Sask., Yukon, Alaska, Ariz., Ark., Calif., Colo., Conn., Del., D.C., Ga., Idaho, Ill., Ind., Iowa, Kans., Ky., Maine, Md., Mass., Mich., Minn., Mo., Mont., Nebr., Nev., N.H., N.J., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.Dak., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wis., Wyo., South America, Eurasia, Africa, Pacific Islands (New Zealand), Australia

Discussion

Ptychostomum pseudotriquetrum is one of the most common and widespread species of Bryaceae, absent only from the subtropics, tropics, and central Pacific Islands. Ovate decurrent leaves, short, stout awn, dense areolation, dioicous sexual condition, and long radiculose stems are characteristic. Ptychostomum bimum is smaller, with much weaker decurrencies, and is synoicous.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

John R. Spence +
(Hedwig) J. R. Spence & H. P. Ramsay ex Holyoak & N. Pedersen +
denticulate +
reduced +
curved +  and straight +
decurrent +
Mnium pseudotriquetrum +  and Bryum pseudotriquetrum +
cucullate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
nutant +, inclined +  and suberect +
long-exserted +
elongate-ovate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
inflated +
not differentiated +
appendiculate +
short-excurrent +
subpercurrent +
rhomboidal;hexagonal +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Ga. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +, Wyo. +, South America +, Eurasia +, Africa +, Pacific Islands (New Zealand) +  and Australia +
low to high elevations (0-4000 m) +
hyaline;pale-yellow +
not adherent +
hyaline +, brown +, orange +  and yellow +
hyaline +  and yellow +
Wet soil, soil over rock, rock, fens +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
polyoicous +, monoicous +, dioicous +  and sexual +
imbricate;variously contorted +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
brick-red +, brown-red +, dull +, yellow-green +, red-green +  and green +
erect;erect-spreading +
ovatelanceolate +  and ovate +
shrunken +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
2-stratose +  and 1-stratose +
revolute entire +  and denticulate +
short-rectangular +  and quadrate +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature Apr–Oct. +
differentiated +
smooth;papillose +
not absent +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
straight +  and flexuose +
elongate +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Illustrated +
green +  and pale-yellow +
germinating +
variable +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
flat +  and weakly concave +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Ptychostomum pseudotriquetrum +
Ptychostomum subg. Cladodium +
species +
lanceolate +
distinct +
not +  and porose +
straight +
specialized +
dioicous +  and sexual +
sinuate +
incrassate +
yellow-green +, red-green +  and green +
large;small to very +