Persicaria bicornis

(Rafinesque) Nieuwland

Amer. Midl. Naturalist 3: 201. 1914.

Common names: Pink smartweed
Endemic
Basionym: Polygonum bicorne Rafinesque Fl. Ludov., 29. 1817
Synonyms: Persicaria longistyla (Small) Small Polygonum longistylum Small
Treatment appears in FNA Volume 5. Treatment on page 589. Mentioned on page 582, 583.

Plants annual, 2–18 dm; roots also rarely arising from basal nodes; rhizomes and stolons absent. Stems ascending to erect, rarely decumbent, branched, ribbed, glabrous or appressed-pubescent to spreading-pubescent distally, stipitate-glandular or, rarely, without stipitate-glands. Leaves: ocrea brownish, cylindric, 6–20 mm, chartaceous, base inflated, margins truncate, eciliate or ciliate with bristles to 1 mm, surface glabrous or scabrous proximally, eglandular; petiole 0.1–1.5 (–2.3) cm, glabrous or appressed-pubescent; blade sometimes with dark triangular or lunate blotch adaxially, linear-lanceolate to ovatelanceolate, 2.3–13 (–18) × (0.4–) 1–2.3 cm, base tapered to cuneate, margins antrorsely scabrous, apex acute to acuminate, faces glabrous or appressed-pubescent along midveins, glandular-punctate abaxially. Inflorescences terminal and axillary, erect, uninterrupted, 8–60 × 10–18 mm; peduncle 8–60 (–70) mm, glabrous or pubescent, usually stipitate-glandular; ocreolae overlapping, margins eciliate or ciliate with bristles to 0.8 mm. Pedicels ascending, 1.5–5 mm. Flowers 2–11 per ocreate fascicle, heterostylous; perianth pink, glabrous, not glandular-punctate, accrescent; tepals 5, connate ca. 1/4–1/3 their length, obovate to elliptic, 3–4.6 mm, veins prominent, not anchor-shaped, margins entire, apex obtuse to rounded; stamens 6–8, included or exserted; anthers pink or red, elliptic; styles 2 (–3), included or exserted, connate at bases. Achenes included or apex exserted, brownish black to black, discoid or, rarely, 3-gonous, 1 side usually slightly concave and other with central hump, (2–) 2.2–2.9 × 2–2.8 (–3) mm, shiny, smooth.


Phenology: Flowering Jun–Oct.
Habitat: Moist, disturbed places, permanent and ephemeral wetlands, ditches, cultivated fields, shorelines of ponds and reservoirs
Elevation: 50-1600 m

Distribution

V5 1201-distribution-map.gif

Ark., Colo., Ill., Iowa, Kans., La., Mo., Nebr., N.Mex., Okla., S.Dak., Tex., Wyo.

Discussion

Persicaria bicornis is a characteristic smartweed of permanent and ephemeral wetlands and moist, disturbed sites in the Great Plains. It often has been included in P. pensylvanica but can be distinguished readily by its heterostylous flowers. The achenes, which usually bear an obscure or prominent hump in the center of one face, also are diagnostic. This hump often ruptures the side of the perianth on fruiting herbarium specimens. Persicaria bicornis also has leaf blades that are on average narrower than are those of P. pensylvanica, and populations exhibit less variation in perianth color. As in P. pensylvanica, flowers with three styles and trigonous achenes are produced very rarely.

Selected References

None.

Lower Taxa

None.

"/2" is not declared as a valid unit of measurement for this property.

... more about "Persicaria bicornis"
dark-brown;black +
exserted +  and included +
spheroidal +, 2-3-gonous +, biconvex +  and discoid +
red +  and pink +
elliptic +  and ovate +
obtuse;rounded +
not +  and swollen +
Harold R. Hinds† +  and Craig C. Freeman +
(Rafinesque) Nieuwland +
tapered +  and cuneate +
Polygonum bicorne +
13 cm130 mm <br />0.13 m <br /> (18 cm180 mm <br />0.18 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2.3 cm23 mm <br />0.023 m <br /> (13 cm130 mm <br />0.13 m <br />) +
linear-lanceolate;ovatelanceolate +
1 cm10 mm <br />0.01 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
lunate +, triangular +  and dark +
foliaceous +  and perfoliate +
scale-like +
0 cm0 mm <br />0 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
just below the soil surface or and above the soil surface +  and spreading +
Pink smartweed +
Ark. +, Colo. +, Ill. +, Iowa +, Kans. +, La. +, Mo. +, Nebr. +, N.Mex. +, Okla. +, S.Dak. +, Tex. +  and Wyo. +
50-1600 m +
glandular-punctate +
appressed-pubescent +  and glabrous +
adnate +, connate +  and distinct +
heterostylous +
disarticulating +  and solid +
decumbent to erect +  and prostrate +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
Moist, disturbed places, permanent and ephemeral wetlands, ditches, cultivated fields, shorelines of ponds and reservoirs +
brownish black +  and black +
0.8 cm8 mm <br />0.008 m <br /> (6 cm60 mm <br />0.06 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
nonmembranous +
sessile +  and petiolate +
deciduous +
entire +  and truncate +
ciliate +  and eciliate +
swollen +
disintegrating +
deciduous +  and persistent +
scabrous +  and variously pubescent +
cylindric +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
coriaceous +
anatropous +, , +  and orthotropous +
accrescent +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
6 cm60 mm <br />0.06 m <br /> (7 cm70 mm <br />0.07 m <br />) +
spreading;erect +
stipitate-glandular;pubescent;glabrous +
0.8 cm8 mm <br />0.008 m <br /> (6 cm60 mm <br />0.06 m <br />) +
not glandular-punctate +
campanulate +
accrescent +
1.5 cm15 mm <br />0.015 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
appressed-pubescent +  and glabrous +
not auriculate +
0.1 cm1 mm <br />0.001 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Flowering Jun–Oct. +
(2-)3(-4)-carpellate +
Amer. Midl. Naturalist +
slender;stout +
0.2 cm2 mm <br />0.002 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.29 cm2.9 mm <br />0.0029 m <br />) +
concave +
0.2 cm2 mm <br />0.002 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
exserted +  and included +
6 +  and 8 +
scandent +  and prostrate +
ascending +  and erect +
, +, stipitate-glandular +, appressed-pubescent +  and spreading-pubescent distally +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
deciduous +
exserted +  and included +
spreading;erect +
scabrous +  and glabrous +
Persicaria longistyla +  and Polygonum longistylum +
Persicaria bicornis +
Persicaria sect. Persicaria +
species +
1/4 +  and 1/3 +
obovate +  and elliptic +
0.3 cm3 mm <br />0.003 m <br /> (0.46 cm4.6 mm <br />0.0046 m <br />) +
coriaceous +
persistent +
membranous +
pubescent +  and glabrous +
not anchor--shaped +
stoloniferous +, rhizomatous +, fibrous-rooted +  and taprooted +
12 +, 11 +  and 10 +