Oenothera cespitosa subsp. marginata

(Nuttall ex Hooker & Arnott) Munz in N. L. Britton et al.

N. Amer. Fl., ser. 2, 5: 101. 1965. (as caespitosa)

Basionym: Oenothera marginata Nuttall ex Hooker & Arnott Bot. Beechey Voy., 342. 1839
Synonyms: Anogra longiflora A. Heller O. cespitosa subsp. eximia (A. Gray) Munz O. cespitosa var. eximia (A. Gray) Munz O. cespitosa var. longiflora (A. Heller) Munz O. cespitosa var. marginata (Nuttall ex Hooker & Arnott) Munz O. eximia A. Gray O. idahoensis Mulford Pachylophus cylindrocarpus A. Nelson P. exiguus (A. Gray) Rydberg P. eximius (A. Gray) Rydberg P. longiflorus (A. Heller) A. Heller P. marginatus (Nuttall ex Hooker & Arnott) Rydberg P. prolatus A. Nelson
Treatment appears in FNA Volume 10.

Herbs caulescent or acaulescent, usually moderately to densely hirsute and glandular puberulent, rarely exclusively glandular puberulent. Stems usually unbranched, rarely with 1–several branches from near base, 10–40 cm. Leaves (2.8–) 10–26 (–36) × (0.6–) 1–3 (–4.5) cm; petiole (3–) 4–11 (–14) cm; blade usually oblanceolate to narrowly elliptic, rarely lanceolate, margins coarsely and irregularly pinnately lobed to dentate or, rarely, serrate. Flowers: floral-tube (41–) 80–140 (–165) mm; sepals (22–) 34–45 (–54) mm; petals fading pink to lavender, (24–) 35–50 (–60) mm; fila­ments (16–) 20–30 (–35) mm, anthers (10–) 12–17 (–20) mm; style (78–) 100–150 (–185) mm. Capsules straight, cylindrical to, sometimes, lanceoloid-cylindrical, slightly asymmetrical, (21–) 25–50 (–68) × 6–8 mm, valve margins with minute to conspicuous tubercles, these sometimes coalesced into a sinuate ridge; pedicel (0–) 1–40 (–55) mm. Seeds usually narrowly to broadly obovoid, rarely suborbicular, 2.2–3.4 × 1.1–2.6 mm, embryo 1/5–1/2 of seed volume, surface appearing longitudinally striate, reticulate under magnification; seed collar large, usually appearing inflated, sealed by a flat membrane which often slightly depressed into seed collar cavity, margin entire or obscurely sinuate distally. 2n = 14.


Phenology: Flowering Mar–Aug.
Habitat: Rocky slopes, cracks in rocks, talus, along gravelly creek beds and arroyos, roadcuts in loose to somewhat compacted soil derived from granite, sandstone, limestone, volcanic cinder, rarely shale, mostly in foothill communities of pinyon-juniper woodlands, big sagebrush scrub, chaparral, grass­lands, openings in ponderosa pine forests.
Elevation: (400–)1200–2300(–3100) m.

Distribution

Ariz., Calif., Colo., Idaho, Nev., N.Mex., Oreg., Tex., Utah, Wash., Wyo., Mexico (Chihuahua), Mexico (Sonora)

Discussion

Selected References

None.

Lower Taxa

None.
1.7 cm17 mm <br />0.017 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
acuminate +, usually acute +  and rounded +
Warren L. Wagner +
(Nuttall ex Hooker & Arnott) Munz in N. L. Britton et al. +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Oenothera marginata +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
indehiscent +  and loculicidal +
subentire +, serrate +, usually coarsely dentate +  and pinnatifid +
lanceolate +, usually oblanceolate +  and narrowly elliptic +
1 +  and several +
recurved +  and erect +
quadrangular +
5 cm50 mm <br />0.05 m <br /> (6.8 cm68 mm <br />0.068 m <br />) +
indehiscent +  and dehiscent +
2.5 cm25 mm <br />0.025 m <br /> (5 cm50 mm <br />0.05 m <br />) +
lanceoloid-cylindrical +  and cylindrical +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
depressed +  and flat +
inflated +  and depressed +
inflated +  and depressed +
Ariz. +, Calif. +, Colo. +, Idaho +, Nev. +, N.Mex. +, Oreg. +, Tex. +, Utah +, Wash. +, Wyo. +, Mexico (Chihuahua) +  and Mexico (Sonora) +
(400–)1200–2300(–3100) m. +
1/5 +  and 1/2 +
whitish +  and green +
exfoliating +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
basifixed +  and versatile +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
14 cm140 mm <br />0.14 m <br /> (16.5 cm165 mm <br />0.165 m <br />) +
hispid +, lanate +  and glabrous +
8 cm80 mm <br />0.08 m <br /> (14 cm140 mm <br />0.14 m <br />) +
zygomorphic +  and actinomorphic +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
unscented +
1 +  and 4 +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Rocky slopes, cracks in rocks, talus, alonRocky slopes, cracks in rocks, talus, along gravelly creek beds and arroyos, roadcuts in loose to somewhat compacted soil derived from granite, sandstone, limestone, volcanic cinder, rarely shale, mostly in foothill communities of pinyon-juniper woodlands, big sagebrush scrub, chaparral, grasslands, openings in ponderosa pine forests.lands, openings in ponderosa pine forests. +
acaulescent +  and caulescent +
perennial +
suffrutescent +
puberulent +  and hirsute +
nodding +  and erect +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
3 cm30 mm <br />0.03 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
deciduous +
10 cm100 mm <br />0.1 m <br /> (26 cm260 mm <br />0.26 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (26 cm260 mm <br />0.26 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
, +, lobed to dentate +  and lobed +
pedicellate +  and sessile +
1 +  and 8 +
4 cm40 mm <br />0.04 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (4 cm40 mm <br />0.04 m <br />) +
5 cm50 mm <br />0.05 m <br /> (6 cm60 mm <br />0.06 m <br />) +
fading pink +  and lavender +
obcordate +  and obovate +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
11 cm110 mm <br />0.11 m <br /> (14 cm140 mm <br />0.14 m <br />) +
4 cm40 mm <br />0.04 m <br /> (11 cm110 mm <br />0.11 m <br />) +
Flowering Mar–Aug. +
N. Amer. Fl., ser. +
sinuate +
basal +  and cauline +
sinuate +
clavate +  and globose +
sinuate +  and entire +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
sculptured +  and smooth +
suborbicular +  and obovoid +
reduced +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
in pairs +  and separating +
4.5 cm45 mm <br />0.045 m <br /> (5.4 cm54 mm <br />0.054 m <br />) +
purple +, pink +, red +  and green +
deciduous +
3.4 cm34 mm <br />0.034 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
decumbent +  and ascending +
10 cm100 mm <br />0.1 m <br /> (40 cm400 mm <br />0.4 m <br />) +
petiolate +  and sessile +
intrapetiolar +
15 cm150 mm <br />0.15 m <br /> (18.5 cm185 mm <br />0.185 m <br />) +
pubescent +  and glabrous +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
roughened +, reticulate +  and papillose +
Anogra longiflora +, O. cespitosa subsp. eximia +, O. cespitosa var. eximia +, O. cespitosa var. longiflora +, O. cespitosa var. marginata +, O. eximia +, O. idahoensis +, Pachylophus cylindrocarpus +, P. exiguus +, P. eximius +, P. longiflorus +, P. marginatus +  and P. prolatus +
Oenothera cespitosa subsp. marginata +
Oenothera cespitosa +
subspecies +
3(-5)-aperturate +
pedicellate +  and sessile +
woody +  and hard +
smooth +  and erose +
papillate +