Mammillaria heyderi var. heyderi

Common names: Heyder’s nipple cactus
Synonyms: Mammillaria applanata Engelmann Mammillaria gummifera var. applanata (Engelmann) L. D. Benson Mammillaria gummifera var. hemispherica (Engelmann) L. D. Benson Mammillaria heyderi var. applanata Engelmann Mammillaria heyderi var. hemispherica
Treatment appears in FNA Volume 4. Treatment on page 256. Mentioned on page 255.

Stems 4–9 × 4–10 (–15) cm; tubercles 9–20 × 3–7 mm. Spines (8–) 14–18 (–27) per areole; radial spines (7–) 13–17 (–26) per areole, abaxial radial spines, needlelike, 6–11 (–16) mm diam., stiff; central spines 0.15–0.35 mm diam. Flowers 1.8–2.3 × 1.8–1.9 cm. 2n = 22.


Phenology: Flowering Mar–Apr; fruiting Aug–Mar.
Habitat: Chihuahuan desert scrub, Tamaulipan thorn scrub, and Edwards Plateau, usually limestone and alluvial substrates
Elevation: 10-1400 m

Distribution

V4 469-distribution-map.gif

N.Mex., Okla., Tex., Mexico (Coahuila), Mexico (Nuevo León), Mexico (Tamaulipas)

Discussion

The spines of Mammillaria heyderi var. heyderi are smaller and usually more numerous than those of the other flat-topped mammillarias in the United States. Individual plants with fewer radial spines, often segregated as var. hemispherica, are frequent in populations of var. heyderi near the Texas coast. Sometimes the central spines are very short. The coarse western var. bullingtoniana has a much narrower range of variation. Reports of M. heyderi var. heyderi from southeastern Arizona and southwestern New Mexico are based on var. bullingtoniana.

Selected References

None.

Lower Taxa

None.
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
cushionlike +
needlelike +  and bristlelike +
circular to linear +
2.5 cm25 mm <br />0.025 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
deciduous +  and persistent +
0.2 cm2 mm <br />0.002 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
plumose +  and hairlike +
0.001 cm0.01 mm <br />1.0e-5 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Allan D. Zimmerman +  and Bruce D. Parfitt +
Muehlenpfordt +
hardened +
triangular +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
curved +  and straight +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.035 cm0.35 mm <br />3.5e-4 m <br />) +
2mm +  and 8mm +
ascending +  and porrect +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.045 cm0.45 mm <br />4.5e-4 m <br />) +
proliferating +
flattened +  and cylindric +
Heyder’s nipple cactus +
not mucilaginous +
N.Mex. +, Okla. +, Tex. +, Mexico (Coahuila) +, Mexico (Nuevo León) +  and Mexico (Tamaulipas) +
10-1400 m +
not separating +
4 cm40 mm <br />0.04 m <br /> (5.2 cm52 mm <br />0.052 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
unconnected +
1.8 cm18 mm <br />0.018 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
rotate +  and campanulate +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (15 cm150 mm <br />0.15 m <br />) +
1.8 cm18 mm <br />0.018 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +
red;crimson;carmine;crimson;carmine;scarlet;red +
indehiscent +
persistent +, long +  and deciduous +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
obovoid +  and clavate +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
4 cm40 mm <br />0.04 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
7.5 cm75 mm <br />0.075 m <br /> (15 cm150 mm <br />0.15 m <br />) +
Chihuahuan desert scrub, Tamaulipan thorn scrub, and Edwards Plateau, usually limestone and alluvial substrates +
1.1 cm11 mm <br />0.011 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
deciduous +  and persistent +
rudimentary +
flat +  and terete +
nearly +  and microscopic +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
tuberculate +
fluted;spheric;depressed-spheric or club-shaped +
brownish +, greenish +, pink +  and tan +
deciduous +  and persistent +
Flowering Mar–Apr +  and fruiting Aug–Mar. +
concave +  and cavernous +
not mucilaginous +
ridgelike +  and nipple--shaped +
Allg. Gartenzeitung +
bristly +  and woolly +
3;2;2;30 +
triangular +
arillate +  and strophiolate +
yellowish +  and reddish-brown +
0.4mm;12mm +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
obovoid;pyriform;obovoid;pyriform;lenticular-reniform;comma--shaped;spheric +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
Illustrated +
1.5 cm15 mm <br />0.015 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
white +  and white-and-brown or brown +
straight +  and curved +
deciduous +  and persistent +
13 (?) +  and 17 (?) +
annulate-ridged +, hairlike +, daggerlike +, subulate +  and acicular +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
hard +  and rigid +
decurrent +
unsegmented +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
purplish +, green +  and gray-green +
4 cm40 mm <br />0.04 m <br /> (9 cm90 mm <br />0.09 m <br />) +
protruding +
flat-topped +  and top--shaped +
4cm;10cm +
red +  and green +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
tuberculate +
Mammillaria applanata +, Mammillaria gummifera var. applanata +, Mammillaria gummifera var. hemispherica +, Mammillaria heyderi var. applanata +  and Mammillaria heyderi var. hemispherica +
adventitious +
epipetric +  and epiphytic +
Mammillaria heyderi var. heyderi +
Mammillaria heyderi +
variety +
fringed +  and entire +
flat +  and concave +
dull +  and glossy +
not confluent +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
9mm +  and 20mm +
cylindric +, conic +  and pyramidal +
3mm +  and 7mm +
straight +
sinuate +
epiphytic;epiphytic;epiphytic +
barrel-shaped +  and spheric +