Ludwigia pilosa

Walter

Fl. Carol., 89. 1788.

Common names: Hairy primrose-willow
Synonyms: Isnardia mollis (Michaux) Poiret I. pilosa (Walter) Kuntze Ludwigia hirsuta Desrousseaux L. mollis Michaux L. rudis Walter
Treatment appears in FNA Volume 10.

Herbs often with prominent aer­enchyma when base sub­merged, forming stolons 30–250 cm, 2–4 mm thick, creeping in mud or floating in water, sometimes bearing flowers and fruits. Stems erect, subterete, densely branched, 40–120 cm, densely hirtellous. Leaves alternate; stipules ovate to lanceolate, 0.2–0.25 × 0.1–0.15 mm, usually obscured by pubes­cence; stolons: petiole 0.2–0.7 cm, blade obovate or elliptic to orbiculate, 0.6–2 × 0.5–1.1 cm, margins with distinct hydathodal teeth, base attenuate, surfaces densely hirtellous to glabrate; stems: petiole 0–0.2 (–1) cm, blade elliptic or lanceolate-elliptic to very narrowly elliptic, 1.5–8 (–10) × 0.3–1.2 (–1.4) cm, base cuneate or attenuate, margins entire with obscure hydathodal glands, apex acute or narrowly acute, surfaces ± densely hirtellous, leaves on branches much reduced; bracts much reduced. Inflorescences usually congested, leafy spikes or racemes, flowers solitary in distal leaf-axils; bracteoles attached 1–2.2 mm distal to base of ovary, linear-lanceolate or narrowly elliptic, 3–6.5 (–7.2) × 0.3–1.5 (–1.7) mm, apex acuminate, surfaces hir­tel­lous. Flowers: sepals ascending with reflexed tips, pale green abaxially, creamy white adaxially, often tinged with pink or red, ovate-deltate, 3.5–5.5 (–6) × 2–4 mm, margins entire, apex elongate-acuminate to subcusp­idate, sur­faces densely hirtellous; petals 0; filaments yellowish, 1.5–2.5 mm, base dilated, anthers 0.6–0.9 (–1.3) × 0.5–0.7 mm; pollen shed in tetrads; ovary obovoid to cupshaped, 2.5–4 × 2.5–4 mm; nectary disc elevated 0.3–0.7 mm on ovary apex, bright-yellow, turning black upon drying, 2–3.6 mm diam., indis­tinctly 4-lobed, densely hirtellous around style base and between lobes; style cream, 1–2 mm, sparsely to densely hirtellous, especially proximally, stigma capitate, 0.3–0.6 × 0.3–0.6 mm, not exserted beyond anthers. Capsules subglobose or, sometimes, oblong-obovoid, subterete or with 4 rounded corners, 3–5 × 3–4.5 mm, hard-walled, irregularly dehiscent, pedicel 0–1 mm. Seeds brown, elliptic-oblong or oblong-ovoid, slightly curved on both ends, 0.5–0.7 × 0.3–0.4 mm, surface cells ± isodiametric. 2n = 32.


Phenology: Flowering Jun–Sep.
Habitat: Roadside ditches, marshes, swales in sandy pine flats, edges of pocosins, peaty bogs, low grassy savannas, swamp forests.
Elevation: 0–300 m.

Distribution

Ala., Fla., Ga., La., Miss., N.C., S.C., Tex., Va.

Discussion

The distribution of Ludwigia pilosa is nearly continuous along the Atlantic and Gulf coastal plains, from extreme southeastern Virginia to northern Florida, and west to Louisiana and southeastern Texas. Disjunct populations occur in northern Alabama and central North Carolina.

Ludwigia pilosa is easily distinguished from most others in sect. Isnardia by being densely hirtellous throughout. Its showy sepals and nectary disc attract multiple insect visitors including ants, bumblebees, honeybees, moths, and wasps (C. I. Peng 1989).

Selected References

None.

Lower Taxa

None.
... more about "Ludwigia pilosa"
prominent +
30 cm300 mm <br />0.3 m <br /> (250 cm2,500 mm <br />2.5 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
3(-5)-aperturate +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
8 cm80 mm <br />0.08 m <br /> (10 cm100 mm <br />0.1 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.5cm;8cm +
elongate-acuminate +, acuminate +, acute +, attenuate +  and cuneate +
0.3cm +  and 1.2cm +
Peter C. Hoch +
Walter +
8 cm80 mm <br />0.08 m <br /> (10 cm100 mm <br />0.1 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (8 cm80 mm <br />0.08 m <br />) +
dilated +, acute +, attenuate +  and cuneate +
0.3cm;1.2cm +
Isnardia +
indehiscent +  and loculicidal +
lanceolate-elliptic +  and elliptic +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
lobed +, toothed +  and entire +
elliptic +  and orbiculate +
reduced +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
reduced +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.72 cm7.2 mm <br />0.0072 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
dark red +  and black +
0.3 cm3 mm <br />0.003 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
elliptic +  and linear-lanceolate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
reduced +
straight +  and slightly curved +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
spreading +  and erect +
with 4 rounded corners +, subterete +, oblong-obovoid +, , +  and subglobose +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Hairy primrose-willow +
Ala. +, Fla. +, Ga. +, La. +, Miss. +, N.C. +, S.C. +, Tex. +  and Va. +
0–300 m. +
hirtellous +
basifixed +  and versatile +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
long-pedicellate +  and subsessile +
straight +  and slightly curved +
by a terminal pore +  and dehiscent +
spreading +  and erect +
cylindric +  and clavate turbinate obpyramidal or globose +
Roadside ditches, marshes, swales in sandy pine flats, edges of pocosins, peaty bogs, low grassy savannas, swamp forests. +
perennial +
glandular-puberulent +, hirtellous +, villous +  and strigillose +
deciduous +
8 cm80 mm <br />0.08 m <br /> (10 cm100 mm <br />0.1 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (8 cm80 mm <br />0.08 m <br />) +
acute +, attenuate +  and cuneate +
0.3cm;1.2cm +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
obovoid +  and cupshaped +
2.5mm +  and 4mm +
numerous +  and 1 +
submerged +
swollen +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
5 +  and 4 +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Jun–Sep. +
Fl. Carol., +
paired +, solitary +  and cluster +
ascending +, decumbent +  and erect +
inconspicuous +
expanded +
basal +  and cauline +
clavate +  and globose +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
50 +  and 400 +
pitted +  and smooth +
oblong-ovoid +  and elliptic-oblong +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
, +, tinged with pink or tinged with red +, creamy white +  and pale green +
persistent +
post-anthesis +
5 +  and 4 +
ovate-deltate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
2 +  and 1 +
unequal +  and subequal +
2 times as many as sepals +
aquatic +, amphibious +  and terrestrial +
lenticular +
2 times as many as sepals +
sprawling +, decumbent +  and prostrate +
hirtellous +
subterete +
40 cm400 mm <br />0.4 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
hemispherical +, capitate +, lobed +  and entire +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
sessile +  and petiolate +
green +  and dark reddish +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
intrapetiolar +
ovate +  and lanceolate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.015 cm0.15 mm <br />1.5e-4 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
aquatic +, amphibious +  and terrestrial +
hirtellous +, densely hirtellous +  and glabrate +
papillate +
Isnardia mollis +, I. pilosa +, Ludwigia hirsuta +, L. mollis +  and L. rudis +
Ludwigia pilosa +
Ludwigia sect. Isnardia +
species +
3(-5)-aperturate +
distinct +