Listera borealis

Morong

Bull. Torrey Bot. Club 20: 31. 1893.

Common names: Northern twayblade
Endemic
Synonyms: Neottia borealis (Morong) Szlachetko Ophrys borealis (Morong) Rydberg
Treatment appears in FNA Volume 26. Treatment on page 590. Mentioned on page 587.

Plants 4–26 cm. Stems green, slender to stout, slightly 4-angled, succulent, glabrous. Leaves: blade green to dark bluish green, elliptic, ovate-elliptic, or lanceolate, 1.3–6 × 0.7–3 cm, apex obtuse to rounded. Inflorescences 5–20-flowered, lax, 20–90 mm; floral bracts lanceolate, ovate, or oblong, 1–3 × 0.5–1 mm, apex obtuse; peduncle and rachis glandular-pubescent. Flowers pale green, yellowish green, or bluish green with veins darker green; pedicel filiform, 3.5–7 mm, glandular-pubescent; sepals and petals strongly reflexed away from lip and column; dorsal sepal elliptic-lanceolate to linear-elliptic, 4–6 × 1.5–2.2 mm, apex obtuse to rounded; lateral sepals linear-elliptic to oblong-lanceolate, falcate, 4.5–7 × 1.4–2.3 mm, apex obtuse to rounded; petals linear to linear-oblong, falcate, 4.5–5 × 0.7–1.5 mm, apex obtuse; lip oblong, slightly narrowed in center, base with broadly rounded and bluntly angled divergent auricles, apex slightly dilated, cleft into 2 oblong or semiorbiculate lobes, with apicule in sinus; disc 3-veined, lateral-veins branched and purplish, base darker green, with ridge in center, thickened along center, 7–12 × 4.2–6.5 mm, margins ciliate; column arcuate, stout, 3–4 × 1 mm. Capsules ellipsoid, 8 × 5 mm. 2n = 56.


Phenology: Flowering Jun–Aug.
Habitat: In moist, rich humus of mossy coniferous or mixed hardwood forests, swamps, often along cold streams fed by melting snow, prefers high acidic soils
Elevation: 1500–3000 m

Distribution

V26 1203-distribution-map.jpg

Alta., B.C., Man., Nfld. and Labr., N.W.T., Nunavut, Ont., Que., Sask., Yukon, Alaska, Colo., Idaho, Mont., Oreg., Utah, Wash., Wyo.

Discussion

In Japan Listera borealis is replaced by L. yatabei Makino, which is nearly identical except for short basal auricles. Listera borealis and L. auriculata are very similar in overall appearance; the ovaries and pedicels in L. borealis are glandular-pubescent, and in L. auriculata they are glabrous.

Leaves occur three in a whorl in Listera borealis forma trifolia Lepage.

Selected References

None.

Lower Taxa

None.
... more about "Listera borealis"
self-pollination +
dilated +, obtuse +  and rounded +
not +  and expanded +
angled +  and rounded +
Lawrence K. Magrath +  and Ronald A. Coleman +
Morong +
darker green +
0.7 cm7 mm <br />0.007 m <br /> (?) +  and 1.2 cm12 mm <br />0.012 m <br /> (?) +
4.2mm +  and 6.5mm +
indehiscent +
not +  and articulate +
conduplicate +  and plicate +
green;green;dark bluish +
1.3 cm13 mm <br />0.013 m <br /> (6 cm60 mm <br />0.06 m <br />) +
lanceolate +, ovate-elliptic +  and elliptic +
0.7 cm7 mm <br />0.007 m <br /> (3 cm30 mm <br />0.03 m <br />) +
8 cm80 mm <br />0.08 m <br /> (?) +
ellipsoid +
5 cm50 mm <br />0.05 m <br /> (?) +
3mm +  and 4mm +
Northern twayblade +
Alta. +, B.C. +, Man. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Colo. +, Idaho +, Mont. +, Oreg. +, Utah +, Wash. +  and Wyo. +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1500–3000 m +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
3-merous +, sessile +  and pedicellate +
bluish green +, yellowish green +  and pale green +
horizontal +  and semierect +
glabrate;glandular-pubescent +
subglobose;ovoid;subglobose;ovoid;ellipsoid +
In moist, rich humus of mossy coniferous or mixed hardwood forests, swamps, often along cold streams fed by melting snow, prefers high acidic soils +
lateral +  and terminal +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
falcate +, linear-elliptic +  and oblong-lanceolate +
subopposite +  and opposite +
cauline +  and basal +
reduced +
not +  and deflexed +
narrowed +  and oblong +
larger +
semiorbiculate +  and oblong +
differing in form and color +
1-many-flowered +  and cymose +
pendent +  and erect +
glandular-pubescent +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
glandular-pubescent +
4.5mm;5mm +
falcate;linear;linear-oblong +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Flowering Jun–Aug. +
Bull. Torrey Bot. Club +
2-100-flowered +
glandular-pubescent +
aerial +  and subterranean +
slender +
minute +
free +, distinct +, adnate +  and connate +
foliaceous +
not +  and alike +
branched +, or +  and simple +
pendent +  and erect +
proliferous +
4-angled +
slender +  and stout +
succulent +
concave +  and convex +
Neottia borealis +  and Ophrys borealis +
Listera borealis +
species +
darker green +
aquatic +, lithophytic +, terrestrial +  and epiphytic +
saprophytic +, photosynthetic +  and mycotrophic +