Lilium pardalinum subsp. pitkinense

(Beane & Vollmer) M. W. Skinner

Novon 12: 255. 2002.

Common names: Pitkin Marsh lily
IllustratedEndemicConservation concern
Basionym: Lilium pitkinense Beane & Vollmer Contr. Dudley Herb. 4: 356, plate 19. 1955
Treatment appears in FNA Volume 26. Treatment on page 190. Mentioned on page 184, 189.

Bulbs branching but less often than in subsp. pardalinum, otherwise similar; scales usually 2-segmented. Stems to 2 m, moderately clonal and usually forming small colonies. Leaves ± evenly distributed along stem, in 2–5 whorls or partial whorls, 3–14 leaves per whorl, ± ascending or horizontal and drooping at the tips, 6.8–22.3 × 1.2–3.6 cm, 4–13 times longer than wide; blade elliptic, sometimes narrowly so, or weakly oblanceolate, margins not undulate. Racemes 1–11-flowered. Flowers not fragrant; sepals and petals reflexed 1/3 along length from base, yellow-orange or orange proximally, darker orange-red to sometimes reddish on distal 1/3–1/2; sepals 4.9–7.1 × 1–1.7 cm; petals 4.9–7.1 × 1.2–2 cm; stamens moderately exserted; filaments moderately spreading, diverging 9°–18° from axis; anthers magenta, 0.6–1.1 cm; pollen red or brown-orange; pistil 3.4–4.6 cm; ovary 1.2–1.9 cm; pedicel 8.5–26.5 cm. Capsules not measured. Seeds not counted.


Phenology: Flowering summer (Jun–Jul).
Habitat: Marshes and adjacent scrub beneath valley oak (Quercus lobata Née)
Elevation: 0–100 m

Discussion

Of conservation concern.

The Pitkin Marsh lily is a local isolate of subsp. pardalinum with small flowers, short stamens, and brighter anthers known only from the vicinity of Sebastopol in Sonoma County, California. Celebrated and once quite common in its limited range, it is now protected by the state of California due to dramatic declines from land clearing for agriculture and housing, hydrological disruption, and depredations by lily fanciers and flower pickers.

The expression of subsp. pitkinense most faithful to the type is extremely rare and found only in Pitkin Marsh. Plants from nearby marshes are somewhat more variable, and some clones begin to approach subsp. pardalinum morphologically; hybrids or plants virtually indistinguishable from the nominate subspecies occur a few miles from Pitkin Marsh. Subspecies pitkinense can be distinguished with some difficulty from subsp. shastense by its darker pollen.

Selected References

None.

Lower Taxa

None.

"narrow" is not a number."wide" is not a number."thicker" is not a number."elongating" is not a number.

0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Mark W. Skinner +
(Beane & Vollmer) M. W. Skinner +
whorled +, opposite +  and alternate +
visible +, exposed +  and hidden +
loculicidal +
dilated +
constricted +
Lilium pitkinense +
reticulate-veined +  and parallel-veined +
paler +  and green and somewhat lighter +
oblanceolate +
rounded +
stained brown +, purplish +, yellowish +  and whitish +
0.2-0.6 times taller than long +
1.4cm +  and 5.1cm +
slanted +  and erect +
chunky +
3.9 cm39 mm <br />0.039 m <br /> (19 cm190 mm <br />0.19 m <br />) +
2.2 cm22 mm <br />0.022 m <br /> (5.7 cm57 mm <br />0.057 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
Pitkin Marsh lily +
0–100 m +
translucent +, pale green +  and variable +
diverging +  and spreading +
connate-coroniform +  and dilated +
sessile +  and pedicellate +
diverging +  and spreading +
not fragrant +
unisexual +  and bisexual +
winged +, 3-valved +  and capsular +
brown;green +
indehiscent +, septicidal +  and loculicidal +
1.5 cm15 mm <br />0.015 m <br /> (7.7 cm77 mm <br />0.077 m <br />) +
oblong-obovate +
membranaceous +  and leathery +
0.8 cm8 mm <br />0.008 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
Marshes and adjacent scrub beneath valley oak (Quercus lobata Née) +
1-28(-35)-flowered +
distributed +
persistent +
6.8 cm68 mm <br />0.068 m <br /> (22.3 cm223 mm <br />0.223 m <br />) +
drooping +, horizontal +  and ascending +
1.2 cm12 mm <br />0.012 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
papillose +  and smooth +
not undulate +
1.2 cm12 mm <br />0.012 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
not articulate +
8.5 cm85 mm <br />0.085 m <br /> (26.5 cm265 mm <br />0.265 m <br />) +
zygomorphic +  and actinomorphic +
s-cap--shaped +
reddish +, darker orange-red to sometimes +, orange +  and yellow-orange +
distinct +
4.9cm;7.1cm +
included +  and strongly exserted +
differentiated +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering summer (Jun–Jul). +
oblong +  and 3-lobed +
3.4 cm34 mm <br />0.034 m <br /> (4.6 cm46 mm <br />0.046 m <br />) +
brown-orange +  and red +
1-11-flowered +
wrinkled +
thinner +  and thick +
2-segmented +
1 cm10 mm <br />0.01 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
starchy +  and fleshy +
abscission +
light-brown +
123 +  and 264 +
verrucose +
wedge +  and flattened +
reddish +, darker orange-red to sometimes +, orange +  and yellow-orange +
distinct +
included;strongly exserted +
ridged +  and clawed +
differentiated +
1 cm10 mm <br />0.01 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
Illustrated +, Endemic +  and Conservation concern +
adnate +  and free +
glabrous +  and glaucous +
0 cm0 mm <br />0 m <br /> (200 cm2,000 mm <br />2 m <br />) +
1 +  and several +
sordid +, pale +  and green +
3 +  and 1 +
Lilium pardalinum subsp. pitkinense +
Lilium pardalinum +
subspecies +
connate +  and distinct +
sepaloid +  and petaloid +
narrower +
3 +  and 14 +