Horkelia fusca var. brownii

(Rydberg) Ertter & Reveal

Novon 17: 319. 2007.

Common names: Shasta horkelia
Endemic
Basionym: Horkelia brownii Rydberg in N. L. Britton et al., N. Amer. Fl. 22: 276. 1908
Treatment appears in FNA Volume 9. Treatment on page 263. Mentioned on page 259, 260, 261, 262.

Stems 1–4.5 dm. Basal leaves grayish, 4–12 (–18) cm; leaflets 3–7 per side, narrowly to broadly cuneate, 5–15 (–17) × 2–8 (–12) mm, usually 1/3–1/2 as wide as long, divided 1/4–1/2 to midrib into 4–6 teeth, surfaces often obscured, ± densely short-villous. Cauline leaves 2–4; leaflets of proximalmost 2–4 per side. Inflorescences grayish or purplish, usually ± congested, sometimes more open, usually comprising less than 1/6 of stem, composed of 5–20 (–40) -flowered glomerules, glandular-hairs usually not red-septate; bracts acuminate-lobed, often ± obscuring pedicels and flowers even at maturity. Flowers: epicalyx bractlets 1.5–2.5 mm; hypanthium 1.5–2.5 × 2.5–4 mm; petals 2–3 (–4.5) mm; filaments (0.2–) 0.6–1 mm, usually longer than wide, anthers 0.3–0.5 mm; styles 1 mm. Achenes 1–1.2 mm. 2n = 28.


Phenology: Flowering summer.
Habitat: Dry meadow edges in conifer woodlands, on pumice or sandy or clay soil
Elevation: 900–2000 m

Discussion

Variety brownii here takes the place of var./subsp. pseudocapitata as used in D. D. Keck (1938) and most floras; this variant was included within var. parviflora by B. Ertter (1993d). The diagnostic feature is the ash gray leaflets, which also tend to be more narrowly cuneate than in other varieties. The variety occurs from Mount Shasta in Siskiyou County east to southwestern Modoc County and southeast to Sierra Valley in Plumas and Sierra counties, California, skirting the range of var. tenella. Morphologically comparable plants occur as far south as the California-Nevada border on the south side of Lake Tahoe, and at least one collection from Nevada (Reno, Hillman s.n., POM) is closest to this variety. Populations from the southeastern end of the range tend to be larger and less ash gray than those in the vicinity of Mount Shasta. Most references to var. brownii (as var./subsp. pseudocapitata misapplied) from Oregon are probably best attributed to the polymorphic var. parviflora, but at least one collection from Klamath Lake (Newberry s.n., GH) fits well in var. brownii.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."/2lengthoftonearlyequaltosepals" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

1mm +  and 1.2mm +
longer than wide +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
pink +  and rose +
less oblanceolate;cuneate +
Barbara Ertter +  and James L. Reveal +
(Rydberg) Ertter & Reveal +
compound +  and simple +
opposite +  and alternate +
20 cm200 mm <br />0.2 m <br /> (40 cm400 mm <br />0.4 m <br />) +
12 cm120 mm <br />0.12 m <br /> (18 cm180 mm <br />0.18 m <br />) +
4 cm40 mm <br />0.04 m <br /> (7 cm70 mm <br />0.07 m <br />) +
4 cm40 mm <br />0.04 m <br /> (20 cm200 mm <br />0.2 m <br />) +
4 cm40 mm <br />0.04 m <br /> (12 cm120 mm <br />0.12 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
flattened +
Horkelia brownii +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
planar +  and cylindric +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
coriaceous +
acuminate-lobed +
reduced +
free +  and distinct +
Shasta horkelia +
open to congested +
Calif. +, Nev. +  and Oreg. +
not +  and aggregated +
900–2000 m +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
usually longer than wide +
flattened +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.4mm;0.6mm +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
5-20(-40)-flowered +
Dry meadow edges in conifer woodlands, on pumice or sandy or clay soil +
1.5mm;2.5mm +
0.1 cm1 mm <br />0.001 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
purplish +  and grayish +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
overlapping +  and separate +
1.5 cm15 mm <br />0.015 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
glabrate +, sparsely +  and more or less densely short-villous or hirsute +
0.2 cm2 mm <br />0.002 m <br /> (?) +  and 0.8 cm8 mm <br />0.008 m <br /> (?) +
oblanceolate +
palmate +  and pinnate +
shallowly toothed +  and deeply divided +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 (?) +  and 4 (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
white +  and pale-pink +
free +  and distinct +
rounded;truncate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
not arillate +
linear;filiform +
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
persistent +
free +  and distinct +
spreading +  and more or less reflexed +
lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
ascending +  and erect +
0 +  and 1/6 +
palmate +  and pinnate +
linear;lanceolate or elliptic +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
1 cm10 mm <br />0.01 m <br /> (?) +
short-villous +
Potentilla douglasii +
Horkelia fusca var. brownii +
Horkelia fusca +
variety +
4 (?) +  and 6 (?) +
oblanceolate +  and obovate +
inconspicuous +
enlarged +
grayish +, reddish +  and green +
hairy +  and inconspicuously conspicuously glandular +