TaxonIllustrator 
FNA25 P85 Eustachys pg 221.jpegEustachys glauca
Eustachys petraea
Karen Klitz
Linda A. Vorobik
Karen Klitz
Linda A. Vorobik

Plants perennial; cespitose, shortly rhizomatous and often stoloniferous. Culms 20-150 cm, erect or decumbent, flattened, glabrous; internodes hollow. Leaves basal and cauline; sheaths open, keeled, strongly compressed, distinctly distichous, equitant; ligules to 0.5 mm, scarious, densely short ciliate; blades flat or folded, erect to spreading, both surfaces usually glabrous, sometimes scabrous or scabridulous. Inflorescences terminal, exceeding the upper leaves, panicles of 1-36 non-disarticulating spikelike branches; branches digitately arranged, axes triquetrous, with spikelets in 2 rows on the abaxial sides of the branches. Spikelets solitary, diverging strongly from the branch axes, laterally compressed, sessile or subsessile, with 2-3 florets; lowest florets bisexual; second florets usually reduced to a stipitate empty lemma, occasionally with a palea, staminate if a third floret present; third florets, if present, sterile and usually rudimentary, stipitate; disarticulation above the glumes. Glumes unequal, 1-veined; lower glumes somewhat smaller than the upper glumes, narrow, acuminate; upper glumes almost as long as the spikelets, flattened, scarious, glabrous, green, pale, or purplish, veins antrorsely scabrous, apices truncate, bilobed, or bifid, often mucronate; lowest lemmas cartilaginous, light to dark-brown, scabridulous distally, 3-veined, unawned, mucronate, or with a single awn, awns to 1.2 mm; paleas equaling or slightly shorter than the lemmas, glabrous, 2-veined, veins keeled, shortly ciliate or scabridulous; anthers 3, deep purple to purple-red. Caryopses 1-1.7 mm, trigonous-ellipsoid, glabrous, translucent when fresh, pale to slightly reddish-purple-tinged, x = 10.

Distribution

Pa., Miss., Tex., La., Calif., Puerto Rico, Virgin Islands, Ala., N.C., S.C., Pacific Islands (Hawaii), Ga., N.J., N.Y., Fla.

Discussion

Eustachys, as treated here, is a genus of approximately 12 species, most of which are native to the Western Hemisphere; four are native to the Flora region and three have been introduced. It is, in many ways, morphologically similar to Chloris, with the placement of a few species being problematic. Molecular data (Alice et al. 2000) support its recognition as a distinct genus, but the relationships between it, Chloris sensu stricto, and Cynodon are not clear.

Key

1 Lateral veins of the lowest lemma in each spikelet usually glabrous, occasionally with a few short, stiff hairs Eustachys glauca
1 Lateral veins of the lowest lemma in each spikelet pubescent. > 2
2 Keels of the lowest lemma in each spikelet glabrous. > 3
3 Spikelets 1.5-2.1 mm long; panicles with 6-15 branches; branches 4-10 cm long, straight, somewhat stiff; leaf blades 5-10 mm wide Eustachys retusa
3 Spikelets 2.4-3 mm long; panicles with 10-36 branches; branches 6-15 cm long, flexible; leaf blades 10-15 mm wide Eustachys distichophylla
2 Keels of the lowest lemma in each spikelet pubescent. > 3
4 Spikelets 1.5-2.5 mm long; lowest lemma in each spikelet mucronate. > 5
5 Lowest lemma in each spikelet dark brown, the lateral veins with appressed hairs shorter than 0.5 mm Eustachys petraea
5 Lowest lemma in each spikelet tawny to reddish-brown, the lateral veins with spreading hairs longer than 0.5 mm Eustachys caribaea
4 Spikelets 2.6-3.7 mm long; lowest lemma in each spikelet awned, awns 0.4-1.2 mm. > 5
6 Panicle branches 1-3; awns of lowest lemma in each spikelet 0.4-0.6 mm long; spikelets 3-3.7 mm long Eustachys floridana
6 Panicle branches (3)4-9; awns of lowest lemma in each spikelet 0.7-1.2 mm long; spikelets 2.6-3 mm long Eustachys neglecta

"decumbent" is not a number.

... more about "Eustachys"
purple +  and purple-red +
bifid +, bilobed +  and truncate +
mucronate +
Cynthia Aulbach +
1 (?) +  and 3 (?) +
0 cm0 mm <br />0 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
triquetrous +
keeled +  and rounded +
pseudopetiolate +  and branching +
not fused +
intravaginal +, extravaginal +, branching +  and basal +
swelling +
membranous +
not pseudopetiolate +
pubescent +  and glabrous +
folded +  and flat +
non-disarticulating +
spikelike +
pale +  and slightly reddish-purple-tinged +
trigonous-ellipsoid +
0.1 cm1 mm <br />0.001 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
triangular +  and dome--shaped +
sometimes longer +
membranous +
decumbent +  and erect +
flattened +
20 cm200 mm <br />0.2 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
not woody +
Pa. +, Miss. +, Tex. +, La. +, Calif. +, Puerto Rico +, Virgin Islands +, Ala. +, N.C. +, S.C. +, Pacific Islands (Hawaii) +, Ga. +, N.J. +, N.Y. +  and Fla. +
not waisted +
capillary +
stipitate +  and staminate +
sometimes longer +
rudimentary +
2 +  and 3 +
round +  and compressed +
reduced +
shorter or longer +
subtending +
unequal +
uncinate +
paniculate +  and racemose +
not absent +
cauline +  and basal +
empty +  and stipitate +
sometimes longer +
0 cm0 mm <br />0 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
inconspicuous +
somewhat smaller +
light +  and dark-brown +
concealed +  and prominent +
2 +  and 1 +
dry +  and fleshy +
spikelike +
alice2000a +  and anderson1974c +
equitant +  and distichous +
compressed +  and keeled +
subsessile +  and sessile +
1 +  and 60 +
compressed +
1 +  and 3 +
compound +  and simple +
2 (?) +  and 3 (?) +
scabridulous +  and scabrous +
Gramineae +
Eustachys +
Poaceae tribe Cynodonteae +
membranous +
purplish +, pale +  and green +
7 (?) +  and 13 (?) +
scabridulous +
stoloniferous +  and rhizomatous +
plant +  and cespitose +
aquatic +  and terrestrial +
dioecious +, monoecious +  and synoecious +