Eriogonum subg. Pterogonum

(H. Gross) Reveal

Sida 3: 82. 1967.

Basionym: Undefined subg. Pterogonum H. Gross Bot. Jahrb. Syst. 49: 239. 1913
Treatment appears in FNA Volume 5. Treatment on page 376. Mentioned on page 221, 222, 223.

Herbs, erect, polycarpic or monocarpic perennials, strigose, silky-pubescent, or glandular-puberulent, occasionally glabrous [tomentose]; taproot woody occasionally chambered. Stems erect, with persistent leaf-bases, silky-pubescent to glandular-puberulent or glabrous; caudex absent or compact, spreading; aerial flowering-stems erect, slender to stout, solid or hollow, sometimes fistulose, not disarticulating in ringlike segments proximally, usually arising directly from root. Leaves persistent or marcescent, basal or basal and cauline, not fasciculate; blade glabrous or strigose to hispid or stipitate-glandular. Inflorescences cymose, open, occasionally with 1 branch of cyme suppressed; bracts 3, connate basally, semileaflike to scalelike. Peduncles ascending to erect, slender. Involucres 1 per node, not appressed to inflorescence branches, turbinate to campanulate; teeth 5, erect. Flowers not attenuate at base, without stipelike base; perianth various shades of white, yellow, red, or purple, glabrous on both surfaces or occasionally pubescent abaxially; tepals connate proximally 1/4–1/3 their length, monomorphic; stamens exserted; filaments pilose proximally or glabrous. Achenes various shades of brown, 3-winged or ridged, 3-gonous, glabrous or pilose [strigose]. Seeds: embryo straight.

Distribution

sw United States, Mexico

Discussion

Species 11 (6 in the flora).

W. J. Hess and J. L. Reveal (1976) subdivided subg. Pterogonum into four sections: sect. Pterogonum (only Eriogonum atrorubens in the flora area), sect. Peregrina W. J. Hess & Reveal (E. greggii), sect. Astra W. J. Hess & Reveal (E. hemipterum, E. hieracifolium, and E. nealleyi), and sect. Alata Bentham (E. alatum). The rest of the species occur in northern Mexico and all belong to sect. Pterogonum: E. ciliatum Torrey ex Bentham, E. clivosum W. J. Hess & Reveal, E. fimbriatum W. J. Hess & Reveal, E. henricksonii Reveal, and E. viscanum W. J. Hess & Reveal.

Key

1 Leaves basal or, if cauline, whorled; stems glandular-puberulent or, if glabrous, stems fistulose > 2
1 Leaves basal; stems strigose to glandular-pubescent or, if glabrous, stems not fistulose > 3
2 Perianths glabrous, purple to red or maroon; cauline leaves absent; stems glabrous, often fistulose; Hidalgo County, New Mexico Eriogonum atrorubens
2 Perianths strigose, yellowish white; cauline leaves whorled; stems glandular-puberulent, not fistulose; Hidalgo County, Texas Eriogonum greggii
3 Plants monocarpic; taproot often chambered; perianths yellow to yellowish green, rarely maroon in anthesis; achenes winged entire length; e Utah, se Wyoming and sw Nebraskato n Arizona, New Mexico, and n and w Texas Eriogonum alatum
3 Plants polycarpic; taproot not chambered; perianths yellow, white to greenish white, reddish purple, or maroon; achenes ribbed or winged only along distal 1/ 2; ec Arizona, wc and s New Mexico, w Texas > 4
4 Inflorescence branches glabrous; perianths white to greenish white, glabrous or sparsely strigose; Coke, Howard, Irion, and Sterling counties, Texas Eriogonum nealleyi
4 Inflorescence branches strigose; perianths yellow to reddish purple or maroon, strigose or pilose; ec Arizona, wc and s New Mexico to sw Texas > 5
5 Perianths reddish purple or maroon, rarely yellow; Brewster County, Texas Eriogonum hemipterum
5 Perianth yellow in anthesis, reddish in fruit; ec Arizona, wc and s New Mexico to w Texas Eriogonum hieracifolium

"/2" is not declared as a valid unit of measurement for this property.

included +  and exserted +
pilose +  and glabrous +
3-gonous +  and ridged +
usually red +  and cream or yellow +
oblong +  and ellipsoid or oval +
James L. Reveal +
(H. Gross) Reveal +
deciduous +  and persistent +
stipelike +
Undefined subg. Pterogonum +
awn-tipped +  and simple +
strigose +  and hispid or stipitate-glandular +
linear;orbiculate +
not awn-tipped +
glandular +, pubescent +  and glabrous +
semileaflike +  and scalelike +
grooved +, disarticulating +  and dichotomous +
not brittle +
puberulent +, pilose-pubescent +, hispid +, sericeous +, floccose +, tomentose +  and lanate +
scabrellous +
ridged +, angled +  and round +
spreading +  and erect +
whorled +, opposite +  and alternate +
sw United States +  and Mexico +
glabrous +  and pilose +
unisexual +, , +  and bisexual +
not attenuate +
not disarticulating +, hollow +  and solid +
decumbent to erect +  and prostrate +
slender;stout +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
open +  and cymose +
capitate +  and subcapitate +
cymelike +, paniclelike +, racemelike +  and spikelike +
connate +  and free +
turbinate +  and campanulate +
nonmembranous +
in whorls +, opposite +, alternate +  and rosulate +
marcescent +
persistent +
cauline +  and basal +
bractlike +
pubescent +  and glabrous +
lobed +, undulate +, crisped +, crenulate +  and entire +
not swollen +
orthotropous +
accrescent +
curved +  and straight +
ascending;erect +
erect to deflexed +
purple;red;purple;red;yellow;white +
pubescent +  and glabrous +
cylindric +  and urceolate +
accrescent +
fleshy +, indurate +  and membranous +
with extrafloral nectaries +  and not articulate +
homostylous +  and 3-carpellate +
hess1976a +
, +  and solid +
slender;stout +
persistent +
decumbent to erect +  and prostrate +
silky-pubescent +  and glandular-puberulent or glabrous +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
consisting of a series +  and tubular +
distinct +
erect;spreading or recurved +
spreading;erect +
slender +  and stout +
Eriogonum subg. Pterogonum +
Eriogonum +
subgenus +
emarginate +  and entire +
1/4 +  and 1/3 +
lobed +  and laciniate +
coriaceous +
persistent +
membranous +
pubescent +  and glabrous +
annual +, biennial +  and perennial +
glabrous +, glandular-puberulent +, silky-pubescent +  and strigose +
monocarpic +  and polycarpic +
3 +, 2 +  and 1 +