Epilobium luteum

Pursh

Fl. Amer. Sept. 1: 259. 1813.

Common names: Yellow willowherb
Treatment appears in FNA Volume 10.

Herbs usually with scaly, sub­terranean soboles, rarely con­densed basal turions. Stems several, ascending to suberect, loosely clumped, subterete, 15–75 cm, simple or sparsely branched distally, subglabrous proximal to inflorescence with densely strigillose lines decur­rent from margins of petioles, mixed strigillose and glandular puberulent distally. Leaves opposite prox­imal to inflorescence, alternate distally, petioles 1–3 mm proximally, subsessile distally; blade ovate or elliptic, 2.5–7.8 × 1.2–3.5 cm, base attenuate, margins den­ticulate with 8–20 low, ± pellucid teeth per side, 4–7 prominent veins per side, apex acute to acuminate, surfaces subglabrous with scattered strigillose hairs on margin and abaxial midrib; bracts much reduced, narrower, more acuminate and denticulate. Inflores­cences nodding, later erect, racemes, congested, simple to sparsely branched, densely mixed strigillose and glandular puberulent. Flowers erect; buds 6–10 × 4–5 mm; pedicel 5–8 mm; floral-tube 1.2–3 × 3–4.6 mm, ring of spreading hairs at mouth inside; sepals greenish cream, lanceolate, 10–12 × 3–3.5 mm, abaxial surface densely glandular puberulent; petals cream to pale-yellow, 12–22 × 9–13 mm, broad apical notch 1.2–2.4 mm; filaments cream, those of longer stamens 13–16.5 mm, those of shorter ones 10–15 mm; anthers yellow, 2.2–3 × 0.7–1.3 mm; ovary 20–35 mm, densely glan­dular puberulent; style cream, 15–22 mm, stigma 4-lobed, 1–2 × 2.8–4.5 mm, lobes spreading to recurved, strongly exserted beyond anthers. Capsules erect, 35–75 mm, surfaces sparsely glandular puberulent; pedicel 10–22 mm. Seeds narrowly fusiform or oblanceoloid, 1–1.2 × 0.4–0.5 mm, chalazal collar 0.06–0.1 mm, surface reticulate; coma persistent, tawny, 6.5–8 mm. 2n = 36.


Phenology: Flowering Jul–Sep.
Habitat: Moist rocky slopes, seeps, banks of lakes, streams, springs, and gravel bars along coastal (boreal) to montane or subalpine areas near snowfields.
Elevation: 0–2200 m.

Distribution

Alta., B.C., Alaska, Calif., Oreg., Wash.

Discussion

Epilobium luteum is one of the most distinctive species in the genus due to its large creamy flowers, a color otherwise known only in the distantly related E. suffruticosum. It appears that E. luteum is most closely related to species of the Alpinae group, sharing not only similarities in perennating habit and structures, but also the derived CC chromosome arrangement (S. R. Seavey and P. H. Raven 1977, 1978).

Because Epilobium luteum is modally outcrossing and commonly pollinated by bees, it sometimes hybri­dizes with several other species of Epilobium when they grow sympatrically. One recurrent hybrid combination is E. luteum × E. ciliatum subsp. glandulosum, first de­scribed and named E. ×treleasianum H. Léveillé, and later E. luteum var. lilacinum L. F. Henderson. Following a suggestion by P. A. Munz (1965) that E. ×treleasianum was a hybrid of that particular parentage, S. R. Seavey (1993) verified the relationship in a series of crossing experiments. Epilobium ×treleasianum occurs over a wide geographical range in Alaska, British Columbia, and Washington, forming repeatedly when the parental species co-occur, and often backcrossing with one or both parents, forming hybrid swarms with varying morphological combinations. It also grows vigorously vegetatively.

Another less common hybrid of Epilobium luteum was described as E. ×pulchrum Suksdorf, and a minor variant as E. ×pulchrum var. albiflorum Suksdorf (S. R. Seavey 1993). The second parent of these hybrids is less obvious, but based on morphological features, it is most likely E. hornemannii, which also grows frequently in sympatry with E. luteum.

Selected References

None.

Lower Taxa

None.

"broad" is not a number."narrower" is not a number.

... more about "Epilobium luteum"
0.22 cm2.2 mm <br />0.0022 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
episepalous +
subequal +  and longer +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
acute +  and acuminate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
Peter C. Hoch +
attenuate +
indehiscent +  and loculicidal +
2.5 cm25 mm <br />0.025 m <br /> (7.8 cm78 mm <br />0.078 m <br />) +
toothed +  and entire +
elliptic +  and ovate +
1.2 cm12 mm <br />0.012 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
erect +  and nodding +
denticulate +  and acuminate +
reduced +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
recurved +  and erect +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
sessile +  and pedicellate +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
3.5 cm35 mm <br />0.035 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
sessile +  and pedicellate +
persistent +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Yellow willowherb +
Alta. +, B.C. +, Alaska +, Calif. +, Oreg. +  and Wash. +
0–2200 m. +
basifixed +  and versatile +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
3mm +  and 4.6mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Moist rocky slopes, seeps, banks of lakes, streams, springs, and gravel bars along coastal (boreal) to montane or subalpine areas near snowfields. +
strigillose +
perennial +  and annual +
suffrutescent +
alternate +  and opposite +
deciduous +
strigillose +
decurrent +
spreading +  and recurved +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
numerous +  and 1 +
multicellular +
1 cm10 mm <br />0.01 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
cream +  and pale-yellow +
12mm +  and 22mm +
obovate +  and obcordate +
0.9 cm9 mm <br />0.009 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Jul–Sep. +
Fl. Amer. Sept. +
simple +  and sparsely branched +
puberulent +  and strigillose +
inflated +
basal +  and cauline +
clavate +  and globose +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
sculptured +  and smooth +
oblanceoloid +  and fusiform +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
greenish cream +
deciduous +
lanceolate +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 times as many as sepals +
branched +  and simple +
mixed +  and clumped +
prostrate +  and decumbent +
ascending +  and suberect +
puberulent +, strigillose +  and subglabrous +
exfoliating +
subterete +
15 cm150 mm <br />0.15 m <br /> (75 cm750 mm <br />0.75 m <br />) +
not woody +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
1.5 cm15 mm <br />0.015 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
aquatic +, amphibious +  and terrestrial +
puberulent +  and subglabrous +
papillose +  and finely reticulate +
constricted +
Epilobium +  and Epilobium sect. Lysimachion +
Epilobium luteum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +