Epilobium densum

Rafinesque

Précis Découv. Somiol. 42. 1814.

Common names: Downy willowherb épilobe dressé
Synonyms: Epilobium strictum Muhlenberg ex Sprengel
Treatment appears in FNA Volume 10.

Herbs with threadlike, sparsely leaved epigeous stolons ter­minating with compact fleshy turions 4–8 × 3–4 mm. Stems erect, simple or loosely clustered, terete, 15–95 cm, often well branched distally, densely vil­lous throughout, sometimes mixed glandular puber­ulent distally, decurrent lines absent. Leaves opposite proximally, alternate and often fasciculate distally, subsessile; blade oblong-lanceolate to sublinear, 2–4.5 × 0.3–0.9 cm, ± exceeding internodes, base cuneate, margins entire or denticulate, 3–6 inconspicuous teeth per side, sometimes revolute, veins apparent on abaxial side, 3–5 per side, apex acute, surfaces villous, especially distally; bracts somewhat reduced. Inflorescences erect racemes, branched or not, densely villous, sometimes mixed glandular puberulent. Flowers erect; buds 2.5–5 × 1.5–1.5 mm; pedicel 3–8 mm; floral-tube 1–1.8 ×1–2 mm, ring of spreading hairs at mouth inside; sepals 2–4.5 × 1–1.3 mm, abaxial surface villous and glan­dular puberulent; petals light to dark-pink, obcordate, 4–6 × 2–3 mm, apical notch 1–1.5 mm; filaments pink, those of longer stamens 2–3.5 mm, those of shorter ones 1–2 mm; anthers cream, 0.5–0.8 × 0.4–0.6 mm; ovary 12–20 mm, densely villous, often mixed glandular puberulent; style cream, 2–3.5 mm, stigma narrowly clavate, entire, 1–1.6 × 0.5–1 mm, surrounded by anthers. Capsules 35–68 mm, short-beaked, surfaces densely villous, sometimes mixed glandular puberulent; pedicel 5–15 mm. Seeds narrowly fusiform to narrowly obovoid, 1.5–2 × 0.5–0.6 mm, chalazal collar inconspic­uous, 0.1 mm, surface low papillose; coma persistent, dingy white, 6–8 mm. 2n = 36.


Phenology: Flowering Jul–Sep.
Habitat: Sphagnum and peat bogs, marshes, seeps, damp pastures.
Elevation: 0–600 m.

Distribution

N.B., N.S., Ont., P.E.I., Que., Conn., Ill., Ind., Iowa, Maine, Md., Mass., Mich., Minn., N.H., N.J., N.Y., Ohio, Pa., R.I., Vt., Va., Wis.

Discussion

Epilobium densum is relatively uncommon but widely distributed within the Great Lakes-St. Lawrence River region in boggy or marshy areas. Within that region, it sometimes grows in close proximity with E. leptophyllum and E. palustre; it is densely pubescent like E. leptophyllum, and although the type of hairs differs markedly on the two species, they sometimes are found mixed on herbarium sheets. Because it is less pubescent, E. palustre is rarely mixed or confused with E. densum. Hybrids among these species occur but are uncommon and generally marked by slightly to moderately reduced seed fertility.

M. L. Fernald (1944d), among others, argued that names published in the Catalogus by Muhlenberg were not validly published. In the case of Epilobium strictum, K. Sprengel considered that Muhlenberg provided enough of a description (upright, soft) to validate the name in 1825. However, Rafinesque had validly published the name E. densum for the same taxon in 1814.

Epilobium molle Torrey is an illegitimate name that pertains here.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium densum"
puberulent +  and villous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
episepalous +
subequal +  and longer +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Peter C. Hoch +
Rafinesque +
cuneate +
indehiscent +  and loculicidal +
2 cm20 mm <br />0.02 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
toothed +  and entire +
oblong-lanceolate +  and sublinear +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
reduced +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and erect +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
sessile +  and pedicellate +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
3.5 cm35 mm <br />0.035 m <br /> (6.8 cm68 mm <br />0.068 m <br />) +
sessile +  and pedicellate +
dingy white +
persistent +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Downy willowherb +  and épilobe dressé +
N.B. +, N.S. +, Ont. +, P.E.I. +, Que. +, Conn. +, Ill. +, Ind. +, Iowa +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, N.H. +, N.J. +, N.Y. +, Ohio +, Pa. +, R.I. +, Vt. +, Va. +  and Wis. +
0–600 m. +
basifixed +  and versatile +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
1mm +  and 2mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Sphagnum and peat bogs, marshes, seeps, damp pastures. +
perennial +  and annual +
suffrutescent +
not +  and branched +
puberulent +  and villous +
alternate +  and opposite +
deciduous +
decurrent +
denticulate +  and entire +
puberulent +  and villous +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
numerous +  and 1 +
multicellular +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
light +  and dark-pink +
4mm +  and 6mm +
obcordate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Jul–Sep. +
Précis Découv. Somiol. +
inflated +
basal +  and cauline +
clavate +  and globose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
sculptured +  and smooth +
narrowly fusiform +  and narrowly obovoid +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
lanceolate +, , +, flushed with red or flushed with cream +  and green +
deciduous +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
2 times as many as sepals +
branched +  and simple +
prostrate +  and decumbent +
glabrous +, villous +, glandular-puberulent +  and strigillose +
exfoliating +
15 cm150 mm <br />0.15 m <br /> (95 cm950 mm <br />0.95 m <br />) +
not woody +
0.1 cm1 mm <br />0.001 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
threadlike +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
aquatic +, amphibious +  and terrestrial +
puberulent +  and villous +
papillose +
constricted +
Epilobium strictum +
Epilobium densum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +
inconspicuous +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +