Epilobium canum subsp. canum

Common names: Hummingbird trumpet
Synonyms: Epilobium canum subsp. angustifolium (D. D. Keck) P. H. Raven E. canum subsp. mexicanum (C. Presl) P. H. Raven Zauschneria californica C. Presl 1831 Z. californica subsp. angustifolia D. D. Keck Z. californica subsp. mexicana (C. Presl) P. H. Raven Z. californica var. microphylla A. Gray ex W. H. Brewer & S. Watson Z. californica var. villosa (Greene) Jepson Z. eastwoodiae Eastwood & Moxley Z. mexicana C. Presl 1831 Z. microphylla (A. Gray ex W. H. Brewer & S. Watson) Moxley Z. velutina Eastwood Z. villosa Greene
Treatment appears in FNA Volume 10.
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Herbs suffruticose or not. Stems 20–110 (–120) cm, ± densely strigillose, densely villous, or both, often mixed short glandular puberulent especially distally, sometimes predominantly short glandular puberulent, rarely glabrate. Leaves alternate and often fasciculate distally; blade usually green to grayish green, rarely silvery-canescent, usually narrowly linear to narrowly lanceolate or elliptic, rarely to narrowly ovate, 0.6–4.5 × 0.1–0.6 (–0.8) cm, margins subentire to remotely serrulate, 4–10 teeth per side, veins usually incon­spicuous, surfaces usually moderately to densely villous and/or strigillose, rarely glabrate. Flowers: floral-tube 17–29 mm; sepals 7–10 mm; petals 8–15 mm. Cap­sules 18–30 mm; subsessile or pedicel 1–3 mm. Seeds 1.5–2.2 × 0.9–1.2 mm. 2n = 30, 60.


Phenology: Flowering (Apr–)Jun–Dec.
Habitat: Dry rocky or stony slopes and ridges, disturbed chaparral, along coastal bluffs, offshore islands, scattered in Sierra Nevada foothills.
Elevation: 0–1000(–2700) m.

Distribution

Calif., Mexico (Baja California)

Discussion

The extremely variable subsp. canum has the low­est elevational range, from sea level to 1000 m through most of its range, and to 2700 m in a few places in the southern Sierra Nevada. It occurs almost exclusively in the Californian Province as defined by R. F. Thorne (1993d), and in the Great Central Valley, Central Western California, and Southwestern California subdivisions of the Jepson Manual (B. G. Baldwinet al. 2012). Specifically, it occurs from north-central California (Lake, Sonoma, and Yolo counties) through the Coast Ranges and along the Pacific Coast south to Baja California, extending to the foothills of the Sierra Nevada (Fresno, Tulare, and Tuolumne counties), with one questionably native disjunct locality in Humboldt County.

In areas where subsp. canum overlaps in range with subsp. latifolium, especially in mountainous areas of southern California (notably Kern, Los Angeles, Monterey, Riverside, and San Bernardino counties), specimens of intermediate morphology occur. Most populations of subsp. canum occur at lower elevations, but where the taxa are sympatric, intermediates can be found. Some specimens from the New York, Pana­mint, and other mountain ranges in the Mojave Desert of San Bernardino County, which extend into Arizona, are treated here as subsp. latifolium, but these have leaves somewhat narrower than typical for that subspecies and approach those of subsp. canum.

Distinctive populations of subsp. canum, consistingof large, shrubby, densely pubescent plants with relatively narrow, fascicled leaves, occur particularly near the coast in southern California. Farther inland and at higher elevations, populations with variable combinations of characters led to a proliferation of names. Although some local series of populations appear quite distinctive, over the entire range there are no combinations of characters that consistently delimit morphological units, nor do any patterns differentiate diploids from tetraploids.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number.

0.27 cm2.7 mm <br />0.0027 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
episepalous +
subequal +  and longer +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Peter C. Hoch +
(Greene) P. H. Raven +
4mm +  and 6.5mm +
cuneate +  and attenuate +
Zauschneria cana +
indehiscent +  and loculicidal +
5 cm50 mm <br />0.05 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
usually green +  and grayish green +
0.6 cm6 mm <br />0.006 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
toothed +  and entire +
silvery-canescent +
usually narrowly linear +  and narrowly lanceolate or elliptic +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
smaller +
1.1 cm11 mm <br />0.011 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
recurved +  and erect +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1.8 cm18 mm <br />0.018 m <br /> (3 cm30 mm <br />0.03 m <br />) +
sessile +  and pedicellate +
curved-ascending +
subclavate +  and fusiform +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
subsessile +  and short-pedicellate +
dingy white +
persistent +
detached +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Hummingbird trumpet +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Calif. +  and Mexico (Baja California) +
0–1000(–2700) m. +
light orange-red +  and white +
basifixed +  and versatile +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
5mm +  and 8mm +
suberect +  and spread +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Dry rocky or stony slopes and ridges, disturbed chaparral, along coastal bluffs, offshore islands, scattered in Sierra Nevada foothills. +
perennial +
not +  and suffruticose +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
deciduous +
decurrent +
subentire +  and remotely serrulate +
villous +  and puberulent +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
numerous +  and 1 +
multicellular +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
white +  and orange-red +
8mm +  and 17mm +
obcordate +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.95 cm9.5 mm <br />0.0095 m <br />) +
Flowering (Apr–)Jun–Dec. +
Ann. Missouri Bot. Gard. +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
inflated +
basal +  and cauline +
clavate +  and globose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
sculptured +  and smooth +
obovoid +
constricted +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
lanceolate +, , +, flushed with red or flushed with cream +  and green +
deciduous +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
2 times as many as sepals +
simple +  and well-branched +
110 cm1,100 mm <br />1.1 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
gray-green +  and green +
prostrate +  and decumbent +
erect +  and ascending +
glabrate +, puberulent +, villous +  and strigillose +
exfoliating +
20 cm200 mm <br />0.2 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
light orange-red +
4.2 cm42 mm <br />0.042 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
aquatic +, amphibious +  and terrestrial +
glabrate +, strigillose +  and villous +
papillose +
constricted +
Epilobium canum subsp. angustifolium +, E. canum subsp. mexicanum +, Zauschneria californica +, Z. californica subsp. angustifolia +, Z. californica subsp. mexicana +, Z. californica var. microphylla +, Z. californica var. villosa +, Z. eastwoodiae +, Z. mexicana +, Z. microphylla +, Z. velutina +  and Z. villosa +
Epilobium canum subsp. canum +
Epilobium canum +
subspecies +
3(-5)-aperturate +
prominent +  and inconspicuous +