Drymocallis glandulosa var. reflexa

(Greene) Ertter

J. Bot. Res. Inst. Texas 1: 43. 2007.

Common names: Greene’s drymocallis or wood beauty
Basionym: Potentilla glandulosa var. reflexa Greene Fl. Francisc., 65. 1891
Synonyms: Drymocallis glandulosa subsp. reflexa (Greene) Soják P. glandulosa subsp. reflexa (Greene) D. D. Keck
Treatment appears in FNA Volume 9. Treatment on page 293. Mentioned on page 287, 292.

Stems 1.5–7.5 dm, base 1.5–3 mm diam. Leaves: basal (3–) 6–20 (–30) cm, leaflet pairs 3 (–4); terminal leaflet ± broadly obovate, (1–) 1.5–5 (–7) × 1–3 cm, teeth usually double, rarely single, 6–16 per side, apex rounded to obtuse; cauline well developed or reduced, leaflet pairs 2–3. Inflorescences (2–) 10–30-flowered, ± leafy or not, usually wide, branch angles (15–) 25–55°. Pedicels 2–10 (proximal to 20) mm. Flowers: epicalyx bractlets narrowly elliptic, 1.5–4 × 0.5–1 (–1.5) mm; sepals reflexed, (3–) 4–6 (–8) mm, apex usually obtuse, sometimes acute; petals reflexed, yellow, narrowly ovate or obovate, 3–4 (–5) × 1.5–3 mm; filaments 0.5–1.5 (–2) mm. Achenes 0.9–1.3 mm. 2n = 14.


Phenology: Flowering May–Aug.
Habitat: Moist, somewhat shaded places, often somewhat disturbed
Elevation: 400–2600 m

Distribution

V9 474-distribution-map.jpg

Calif., Nev., Oreg., Mexico (Baja California)

Discussion

At its morphologic extreme, var. reflexa is distinctive in its small, elliptic-oblanceolate, reflexed yellow petals, and widely branched inflorescences; however, in northern California and southern Oregon, var. reflexa and var. glandulosa completely intergrade. As circumscribed here, var. reflexa occurs at low to middle elevations in the mountains from northern Baja California, Mexico, and the Peninsular and Transverse ranges of southern California, through the Sierra Nevada to the mountains of northern California, barely entering adjacent Nevada and Oregon. Some disjunct collections from central Idaho approach this variety; their optimal disposition remains unclear.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."wide" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

reddish-brown +
0.9mm +  and 1.3mm +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
acute;obtuse;rounded;obtuse +
Barbara Ertter +
(Greene) Ertter +
compound +  and simple +
opposite +  and alternate +
20 cm200 mm <br />0.2 m <br /> (30 cm300 mm <br />0.3 m <br />) +
6 cm60 mm <br />0.06 m <br /> (20 cm200 mm <br />0.2 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Potentilla glandulosa var. reflexa +
oblanceolate-oblong +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
Greene’s drymocallis or wood beauty +
Calif. +, Nev. +, Oreg. +  and Mexico (Baja California) +
not +  and aggregated +
400–2600 m +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
linear-lanceolate +  and elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
less;10;80 +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Moist, somewhat shaded places, often somewhat disturbed +
not +, leafy +  and (2-)10-30-flowered +
1/10 +  and 1/5-3/4 +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
overlapping +  and not confluent +
elliptic-ovate +  and flabellate or rhombic +
toothed +, or +, flat +, entire +  and more or less toothed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not overlapping +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0 (?) +  and 4 (?) +
free +  and distinct +
obovate;ovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering May–Aug. +
adnate +, free +, connate +  and distinct +
J. Bot. Res. Inst. Texas +
not arillate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
persistent +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Illustrated +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
1 +  and 20 +
palmate +  and pinnate +
narrowly;broadly-triangular-ovate +
deciduous +
distinct +
fusiform +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Drymocallis glandulosa subsp. reflexa +  and P. glandulosa subsp. reflexa +
Drymocallis glandulosa var. reflexa +
Drymocallis glandulosa +
variety +
5 cm50 mm <br />0.05 m <br /> (7 cm70 mm <br />0.07 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (5 cm50 mm <br />0.05 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
horseshoe--shaped +
mucronate +
6 (?) +  and 16 (?) +
inconspicuous +
hemispheric +  and conic +
enlarged +
peglike-glandular +  and septate-glandular +