Cyperus retroflexus

Buckley

Proc. Acad. Nat. Sci. Philadelphia. 14: 9. 1862.

Illustrated
Synonyms: Cyperus uniflorus var. floribundus Kükenthal Cyperus uniflorus var. retroflexus (Buckley) Kükenthal Mariscus uniflorus
Treatment appears in FNA Volume 23. Mentioned on page 186.

Herbs, perennial, cespitose, rhizomatous. Culms trigonous, basally tuberous-thickened, (5–) 15–40 (–80) cm × 1–2.5 mm, glabrous. Leaves flat to V-shaped, (5–) 10–30 (–60) cm × (0.5–) 1–3 (–4) mm. Inflorescences: spike 1, loosely to densely ovoid to cylindric, 8–36 × 8–20 mm; rays 3–6 (–9), (5–) 10–32 cm; bracts 3–6, horizontal to ascending at 30°, V-shaped, (1–) 3–15 (–30) cm × 0.5–4.5 mm; rachilla deciduous, wings 0.5–0.7 mm wide. Spikelets (5–) 20–35 (–65), linear to oblong (without narrowed basal portion), quadrangular; (2.2–) 4–10 (–18) × 0.7–1.1 mm; floral scales persistent, 1–5, appressed, pale greenish white laterally, stramineous with reddish dots, or deep red, greenish or light-brown medially, 3–6-ribbed laterally, narrowly ovate to oblongelliptic, 2.6–3.9 × 1.6–2 mm, apex rounded to ± acute, entire, mucronulate; distal scale with glabrous midrib and mucro 0.1–0.3 (–0.5) mm; sterile terminal scale of spikelet only 0.4–0.6 mm wide, involute, frequently uncinate. Flowers: anthers (0.3–) 0.4–0.6 mm; styles 0.4–0.5 mm; stigmas 0.5–0.9 mm. Achenes light-brown to reddish-brown, sessile or slightly stipitate, ellipsoid, (1.8–) 2–2.4 × 0.6–0.8 (–0.9) mm, apex slightly apiculate, surfaces puncticulate.


Phenology: Fruiting mid summer–early fall (Jul–Sep).
Habitat: Damp, disturbed soils, croplands
Elevation: 0–1000 m

Distribution

V23 321-distribution-map.jpg

Ala., Ariz., Ark., Miss., Mo., N.Mex., Okla., Tex., Mexico

Discussion

Cyperus retroflexus is distinguished from other species of Cyperus with deciduous spikelets by the involute, sterile terminal scale of the spikelet. In plants with cuspidate scales, the terminal scale is usually somewhat recurved, giving the appearance of a hook at the end of the spikelet.

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Cyperus retroflexus"
stipitate +  and sessile +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
light-brown +  and reddish-brown +
0.2 cm2 mm <br />0.002 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
ellipsoid +
0.6mm;0.8mm +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
apiculate +  and entire +
mucronulate;rounded;more or less acute +
Gordon C. Tucker* +, Brian G. Marcks* +  and J. Richard Carter * +
Buckley +
persistent +  and deciduous +
decurrent +
w--shaped +  and keeled +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
3 cm30 mm <br />0.03 m <br /> (15 cm150 mm <br />0.15 m <br />) +
horizontal +  and ascending +
v--shaped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
parallel +  and divergent +
terete +, rolled +  and plicate +
40 cm400 mm <br />0.4 m <br /> (80 cm800 mm <br />0.8 m <br />) +
15 cm150 mm <br />0.15 m <br /> (40 cm400 mm <br />0.4 m <br />) +
scabridulous +
trigonous +
tuberous-thickened +
1mm;2.5mm +
Ala. +, Ariz. +, Ark. +, Miss. +, Mo. +, N.Mex. +, Okla. +, Tex. +  and Mexico +
0–1000 m +
light-brown +, greenish +, red +, deep +, punct +, stramineous with reddish dots +, list +, coloration +, white +  and pale greenish +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.39 cm3.9 mm <br />0.0039 m <br />) +
narrowly ovate +  and oblongelliptic +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
hypogynous +  and subtending +
biconvex +  and trigonous +
Damp, disturbed soils, croplands +
subumbellate +  and capitate +
spreading +  and erect +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
30 cm300 mm <br />0.3 m <br /> (60 cm600 mm <br />0.6 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
10 cm100 mm <br />0.1 m <br /> (?) +  and 30 cm300 mm <br />0.3 m <br /> (?) +
flat +  and v-shaped +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting mid summer–early fall (Jul–Sep). +
2-3(-4)-carpellate +
antrorse +  and extrorse +
3 (?) +  and 1 (?) +
Proc. Acad. Nat. Sci. Philadelphia. +
continuous +, articulate +, winged +  and wingless +
conspicuous +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
unequal +
3 +  and 6 +
10 cm100 mm <br />0.1 m <br /> (32 cm320 mm <br />0.32 m <br />) +
adventitious +
basal +  and proximal +
cylindric;compressed +
cylindric +
Illustrated +
0.8 cm8 mm <br />0.008 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
ovoid;cylindric +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
20 +  and 35 +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
quadrangular;linear;oblong +
0.7mm;1.1mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
2-3-fid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Cyperus uniflorus var. floribundus +, Cyperus uniflorus var. retroflexus +  and Mariscus uniflorus +
Cyperus retroflexus +
Cyperus subg. Cyperus +
species +
herb +  and cespitose +
0.5mm +  and 0.7mm +