Circaea alpina subsp. pacifica

(Ascherson & Magnus) P. H. Raven

Canad. J. Bot. 43: 1396. 1965.

Basionym: Circaea pacifica Ascherson & Magnus Bot. Zeitung (Berlin) 29: 392. 1871
Synonyms: C. alpina var. pacifica (Ascherson & Magnus) M. E. Jones
Treatment appears in FNA Volume 10.

Stems firm, terete, remaining mostly unflattened after pressing, pubescent with at least a few recurved, falcate hairs. Leaves: petiole 1.5–5 cm; blade 3–7.5 (–11) × 2.5–5.5 (–8) cm, margins subentire to minutely denticulate, base usually rounded to subcordate, rarely cordate, apex acute to short acuminate. Inflorescences usually densely, less often sparsely, glandular puberulent. Flowers clustered at apex of raceme, opening before elongation of raceme axis; floral-tube 0.3–0.6 mm; apical notch of petal 1/4–1/3 length of petal; pedicel and mature fruit combined length 3.5–6.5 mm. 2n = 22.


Phenology: Flowering summer.
Habitat: Cool, temperate deciduous and mixed forests, forest margins, along streams.
Elevation: (0–)200–2700(–2900) m.

Distribution

Cool, temperate deciduous and mixed forests, forest margins, along streams, Alta., B.C., Alaska, Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo.

Discussion

Subspecies pacifica and subsp. alpina are easily separated by the stems with at least a few hairs and by the minutely denticulate to nearly entire leaf blade margins in subsp. pacifica. The stems of subsp. pacifica remain unflattened in herbarium specimens, whereas the stems of subsp. alpina are flattened and appear to be very narrowly winged after pressing. Subspecies pacifica is usually larger and more robust than subsp. alpina and is similar to some of the Asian subspecies of Circaea alpina in its somewhat thicker, deeper green leaves and less delicate stems. Plants in the areas where the two subspecies are sympatric are sometimes difficult to assign to one subspecies or the other, the few minute hairs on the stem being the most reliable distinguishing feature.

Selected References

None.

Lower Taxa

None.
acute +  and short acuminate +
David E. Boufford +
(Ascherson & Magnus) P. H. Raven +
cordate +, usually rounded +  and subcordate +
Circaea pacifica +
indehiscent +  and loculicidal +
7.5 cm75 mm <br />0.075 m <br /> (11 cm110 mm <br />0.11 m <br />) +
5.5 cm55 mm <br />0.055 m <br /> (8 cm80 mm <br />0.08 m <br />) +
3 cm30 mm <br />0.03 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
dentate +  and prominently dentate +
lobed +, toothed +  and entire +
usually ovate +  and broadly ovate +
2.5 cm25 mm <br />0.025 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
reflexed +  and spreading +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0 cm0 mm <br />0 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
ringlike +  and cylindrical +
Cool +, temperate deciduous and mixed forests +, forest margins +, along streams +, Alta. +, B.C. +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +  and Wyo. +
(0–)200–2700(–2900) m. +
basifixed +  and versatile +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
indehiscent +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
reflexed +  and spreading +
obovoid +  and globose +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.78 cm7.8 mm <br />0.0078 m <br />) +
Cool, temperate deciduous and mixed forests, forest margins, along streams. +
falcate +
perennial +
puberulent +, glandular +, glabrous +  and pubescent +
branched +  and simple +
12 cm120 mm <br />0.12 m <br /> (17 cm170 mm <br />0.17 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (12 cm120 mm <br />0.12 m <br />) +
deciduous +
subentire +  and minutely denticulate +
numerous +  and 1 +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
ascending +  and erect +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.78 cm7.8 mm <br />0.0078 m <br />) +
1/4 length of petal +  and 1/3 length of petal +
obdeltate +  and obovate or broadly obovate +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Flowering summer. +
Canad. J. Bot. +
rounded +
basal +  and cauline +
several +  and 1 +
clavate +  and globose +
11 +, 2 +  and 1 +
sculptured +  and smooth +
ellipsoid +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
purple-tinged +, pink +  and white +
deciduous +
reflexed +  and spreading +
ovate +  and broadly ovate +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
aquatic +, amphibious +  and terrestrial +
2 times as many or as many as sepals +
branched +  and unbranched +
prostrate +  and decumbent +
3 cm30 mm <br />0.03 m <br /> (50 cm500 mm <br />0.5 m <br />) +
obpyramidal +  and bilobed +
intrapetiolar +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
aquatic +, amphibious +  and terrestrial +
with prominent longitudinal grooves +  and smooth +
papillate +
C. alpina var. pacifica +
Circaea alpina subsp. pacifica +
Circaea alpina +
subspecies +
3(-5)-aperturate +