Camissoniopsis cheiranthifolia

(Hornemann ex Sprengel) W. L. Wagner & Hoch

Syst. Bot. Monogr. 83: 204. 2007.

Basionym: Oenothera cheiranthifolia Hornemann ex Sprengel Syst. Veg. 2: 228. 1825
Synonyms: Agassizia cheiranthifolia (Hornemann ex Sprengel) Spach Camissonia cheiranthifolia (Hornemann ex Sprengel) Raimann Holostigma cheiranthifolium (Hornemann ex Sprengel) Spach Sphaerostigma cheiranthifolium (Hornemann ex Sprengel) Fischer & C. A. Meyer
Treatment appears in FNA Volume 10.

Herbs short-lived perennial, sometimes woody at base, usually densely strigillose throughout, rarely glabrous, also villous distally. Stems prostrate, decumbent, or ascending from base, to 60 (–130) cm. Leaves 0.5–5 × 0.3–2.2 cm; petiole 0–1.5 (–2.5) cm, distal ones to 1 cm; blade narrowly ovate, base attenuate, cuneate, or cordate, margins sparsely serrulate, apex acute. Flow­ers opening near sunrise; floral-tube 2.1–8.5 mm; sepals 4–11.5 mm; petals yellow, often red-dotted near base, 6–20 mm; episepalous filaments 2.8–8 mm, epipetalous filaments 1.5–6 mm, anthers 1–3 mm, less than 5% of pollen-grains 4-pored or 5-pored; style 6–23 mm, stigma surrounded by or exserted beyond anthers at anthesis. Capsules often coiled in 1–2 spirals, 4-angled, 10–25 × 2–2.5 mm. Seeds 1.2–1.3 mm.

Distribution

w United States, nw Mexico

Discussion

Subspecies 2 (2 in the flora).

Camissoniopsis cheiranthifolia occurs on slopes and dunes along the immediate coast and on islands from Coos Bay, Curry County, Oregon, to the vicinity of San Quintín, Baja California; it is also known from the east shore of San Francisco Bay and locally on sand dunes along the lower Sacramento River, California, 0–100 m. P. H. Raven (1969) determined C. cheiranthifolia to be self-incompatible (some populations in subsp. suffruticosa) or self-compatible (both subspecies) and apparently pollinated by oligolectic bees of Andrena subg. Onagrandrena (Raven); Raven subdivided the species into two intergrading subspecies.

Selected References

None.

Key

1 Plants primarily herbaceous; petals 6–11 mm; style 6–9 mm; stigma usually surrounded by anthers. Camissoniopsis cheiranthifolia subsp. cheiranthifolia
1 Plants woody at base; petals (10–)12–20 mm; style 13–23 mm; stigma exserted beyond anthers at anthesis. Camissoniopsis cheiranthifolia subsp. suffruticosa
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Warren L. Wagner +
(Hornemann ex Sprengel) W. L. Wagner & Hoch +
cordate +, cuneate +  and attenuate +
Oenothera cheiranthifolia +
indehiscent +  and loculicidal +
serrulate +, denticulate +  and dentate +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1-5 times or straight narrowly cylindrical and thickened +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
w United States +  and nw Mexico +
white +  and reddish-brown +
exfoliating +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
basifixed +  and versatile +
0.21 cm2.1 mm <br />0.0021 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
actinomorphic +
not swollen +, 4-angled +, curled +  and contorted +
1-5 times or straight narrowly cylindrical and thickened +
perennial +  and short-lived +
villous +, glabrous +  and strigillose +
deciduous +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
numerous +  and 1 +
red-dotted +  and yellow +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Syst. Bot. Monogr. +
basal +  and cauline +
clavate +  and globose +
7 +  and numerous +
sculptured +  and smooth +
obovoid +  and flattened +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
purple +, pink +, red +  and green +
deciduous +
0.4 cm4 mm <br />0.004 m <br /> (1.15 cm11.5 mm <br />0.0115 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
2 times as many or as many as sepals +
60 cm600 mm <br />0.6 m <br /> (130 cm1,300 mm <br />1.3 m <br />) +
ascending +  and decumbent +
0 cm0 mm <br />0 m <br /> (60 cm600 mm <br />0.6 m <br />) +
subcapitate +  and subglobose +
petiolate +  and sessile +
intrapetiolar +
pubescent +  and glabrous +
0.6 cm6 mm <br />0.006 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
aquatic +, amphibious +  and terrestrial +
non-papillate +
Agassizia cheiranthifolia +, Camissonia cheiranthifolia +, Holostigma cheiranthifolium +  and Sphaerostigma cheiranthifolium +
Camissoniopsis cheiranthifolia +
Camissoniopsis +
species +
3(-5)-aperturate +
smooth +  and erose +
papillate +