Asteraceae (tribe Heliantheae) subtribe Guardiolinae

H. Robinson

Phytologia 41: 41. 1978.

Treatment appears in FNA Volume 21. Treatment on page 41. Mentioned on page 7.

Perennials or subshrubs, (10–) 30–100 [–200] cm. Leaves cauline; opposite; ± petiolate; blades deltate or rounded-deltate to ovate [lanceolate to linear] (bases often hastate), margins usually dentate, faces glabrous or glabrate. Heads radiate, in corymbiform arrays [borne singly]. Calyculi 0. Involucres narrowly cylindric [campanulate]. Phyllaries persistent, 3–5 in 1 series (distinct, oblong or lanceolate to linear, subequal, herbaceous, conduplicate, streaked with 8–12+ translucent nerves, margins often membranous). Receptacles convex, paleate (paleae similar to phyllaries, membranous, conduplicate, each enfolding a floret). Ray-florets 1–5, pistillate, fertile; corollas whitish (tubes often longer than laminae). Disc-florets 3–20+, functionally staminate; corollas whitish, tubes longer than abruptly dilated, campanulate to funnelform throats, lobes 5, lanceolate to lance-linear; anther thecae green (filaments hairy); stigmatic papillae 0 or vestigial. Cypselae ± ellipsoid [fusiform to clavate], ± obcompressed, smooth or fine-ribbed, glabrous (bases carunculate); pappi 0 [5–6, erose scales].

Distribution

w United States, Mexico

Discussion

Genus 1, species 10 (1 species in the flora).

Guardiola has been included in Melampodiinae (G. Bentham 1873) and in Coreopsidinae (T. F. Stuessy 1973). According to H. Robinson (1981), the “glands on the anther appendages” in Guardiola “are distinct and massive, unlike...other Heliantheae.”

Selected References

None.

Lower Taxa

not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
Theodore M. Barkley† +, Luc Brouillet +  and John L. Strother +
H. Robinson +
decurrent +
compound +  and simple +
rounded-deltate;ovate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
fine-ribbed +  and smooth +
obcompressed +  and ellipsoid +
fertile +  and bisexual +
w United States +  and Mexico +
glabrate +  and glabrous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
lanceolate +  and lance-linear +
2-carpellate +
persistent +
bearing subulate enations +, hairy +  and bristly +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Asteraceae (tribe Heliantheae) subtribe Guardiolinae +
Asteraceae tribe Heliantheae +
subtribe +
longer than abruptly dilated +
campanulate +
100 cm1,000 mm <br />1 m <br /> (200 cm2,000 mm <br />2 m <br />) +
30 cm300 mm <br />0.3 m <br /> (100 cm1,000 mm <br />1 m <br />) +
100 cm1,000 mm <br />1 m <br /> (200 cm2,000 mm <br />2 m <br />) +