Harmonia

B. G. Baldwin

Novon 9: 463. 1999.

Etymology: For Harvey Monroe Hall, 1874–1932, Californian botanist
Treatment appears in FNA Volume 21. Treatment on page 297. Mentioned on page 255, 298, 300.
 TaxonIllustrator 
FNA21 P47 Harmonia hallii.jpegHarmonia hallii
Anisocarpus madioides
Jensia yosemitana
Marjorie C. Leggitt
Yevonn Wilson-Ramsey
Barbara Alongi

Annuals, 5–40 cm. Stems erect. Leaves mostly cauline; proximal opposite, distal alternate; sessile; blades linear, margins entire or toothed, faces usually hirsute, sometimes minutely stipitate-glandular as well (glands usually black, sometimes yellowish). Heads radiate, borne singly or in loose, ± umbelliform to corymbiform arrays. Peduncular bracts: pit-glands, tack-glands, and spines 0. Involucres obovoid to obconic, 2–5+ mm diam. Phyllaries 3–8 in 1 series (lanceolate to oblanceolate, herbaceous, strongly conduplicate, each wholly enveloping a ray ovary). Receptacles flat to convex, glabrous or setulose, paleate (paleae falling, in 1 series between rays and discs, distinct or weakly connate, phyllary-like, more scarious). Ray-florets usually 3–8, pistillate, fertile; corollas bright-yellow (laminae flabelliform to obovate). Disc-florets 7–30, bisexual and fertile or functionally staminate (sometimes in same head); corollas bright-yellow, tubes shorter than narrowly funnelform throats, lobes 5, deltate (anthers yellowish to brownish; styles glabrous proximal to branches). Ray cypselae (black) terete to ± compressed, weakly arcuate, gibbous or not, beaked or beakless, glabrous; pappi 0 or of 3–12 lanceolate to subulate, fimbrillate to plumose scales. Disc cypselae (black) ± terete, ± clavate, glabrous or hairy; pappi of 7–11 lance-attenuate, linear, oblong, quadrate, or subulate, fimbriate, fimbrillate, or plumose scales. x = 9.

Discussion

Species 5 (5 in the flora).

Members of Harmonia occur in mountains of northwestern California, as far south as the northern San Francisco Bay area. All but H. nutans are known only from serpentine exposures and are probably descended from a common, serpentine-endemic ancestor (B. G. Baldwin 2001). Harmonia has been treated in Madia, which is more closely related to Carlquistia than to Harmonia (B. G. Baldwin 1996). See Baldwin (2001) for discussion of phylogeny within Harmonia.

Key

1 Heads usually reflexed in bud and fruit; ray pappi 0; disc pappi of lance-attenuate, fimbrillatescales 2–3.7 mm Harmonia nutans
1 Heads usually erect in bud and fruit; ray pappi 0.2–1.5 mm; disc pappi of lanceolate, linear, oblong, quadrate, or subulate, fimbriate or plumose scales 0.2–3.5 mm > 2
2 Leaves ± evenly distributed on stems; ray cypselae strongly gibbous (bowed abaxially), beaked (beaks 0.4–0.5 mm); disc florets functionally staminate Harmonia doris-nilesiae
2 Leaves unevenly distributed, mostly along primary (central) stems and immediately proximal to branches supporting heads; ray cypselae slightly or not gibbous, beakless (pappi elevated adaxially 0.1–0.2 mm in H. stebbinsii); disc florets (some or all) bisexual, fertile > 3
3 Phyllaries densely white-villous near folded edges; ray cypselae slightly gibbous; disc pappi of linear-attenuate to subulate, plumose scales 1.2–3.5 mm Harmonia stebbinsii
3 Phyllaries hirsute and/or hirtellous near folded edges; ray cypselae not gibbous; disc pappi of lanceolate, linear, oblong, or quadrate, fimbriate scales 0.2–0.8 mm > 4
4 Proximal, unbranched portions of primary (central) stems usually shorter than branches supporting heads (distal leaves of primary stems densely congested); disc pappi of oblong or quadrate scales 0.2–0.5 mm Harmonia hallii
4 Proximal, unbranched portions of primary (central) stems usually longer than branches supporting heads (distal leaves of primary stems not densely congested);disc pappi of lanceolate to linear scales 0.6–0.8 mm Harmonia guggolziorum
... more about "Harmonia"
not tailed +, rounded +  and obtuse +
pale +  and dark +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
less umbelliform +  and corymbiform +
indeterminate +  and determinate +
Bruce G. Baldwin +  and John L. Strother +
B. G. Baldwin +
decurrent +
compound +  and simple +
linear to linear-elliptic +
lobed;usually spatulate;oblanceolate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
bright-yellow +
hairy +  and glabrous +
fusiform +, clavate +, obpyramidal +, terete +, compressed +  and obcompressed +
fertile +  and bisexual +
hairy +  and glabrous +
clavate +  and terete +
fertile +  and bisexual +
For Harvey Monroe Hall, 1874–1932, Californian botanist +
stipitate-glandular +  and hirsute +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
in loose , more or less umbelliform to corymbiform arrays +  and singly +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
obovoid;obconic +
petiolate +  and sessile +
alternate +  and opposite +
deltate +
dentate to pinnatifid or palmatifid +  and entire +
toothed +  and entire +
2-carpellate +
persistent +
of 3 +  and 12 lanceolate to subulate fimbrillate to plumose , scales +
beakless +  and beaked +
not +, gibbous +, terete +  and more or less compressed +
setulose +  and glabrous +
flat;convex +
baldwin2001b +
7 +  and 11 +
plumose +, fimbrillate +, fimbriate +, subulate +, quadrate +, oblong +, linear +, lance-attenuate +, lanceolate +  and subulate fimbrillate +
exalbuminous +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Harmonia +
Asteraceae (tribe Heliantheae) subtribe Madiinae +
cylindric +
toothed +  and entire +
5 cm50 mm <br />0.05 m <br /> (40 cm400 mm <br />0.4 m <br />) +
shrub +  and subshrub +