Chaenactis glabriuscula

de Candolle in A. P. de Candolle and A. L. P. P. de Candolle

in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 659. 1836.

Common names: Yellow pincushion
Treatment appears in FNA Volume 21. Treatment on page 411. Mentioned on page 409.

Plants 6–60 cm; proximal indument grayish to whitish, arachnoid to densely lanuginose, or glabrescent. Stems mostly 1–5 (–12; sometimes ± horizontal); branches proximal and, often, distal. Leaves basal (often withering) and cauline, 1–10 cm; largest blades linear or ± elliptic, plane to 3-dimensional, succulent or not, (0–) 1–2-pinnately lobed; primary lobes 1–7 pairs, remote to ± congested, ultimate lobes ± plane, involute, twisted, and/or terete. Heads (± radiant) mostly 1–20+ per stem. Peduncles 1–20 (–30) cm, distally stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent. Involucres ± hemispheric to obconic or broadly cylindric. Phyllaries: longest 4.5–10 mm; outer stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent in fruit, apices erect, blunt, ± rigid. Florets: corollas bright to dark yellow, 4–8 mm (inner); peripheral corollas spreading, ± zygomorphic, enlarged. Cypselae 3–9 mm (± terete); pappi of (1–) 4 scales in 1 series, or of (5–) 8 scales in 2, abruptly unequal series, longest scales (1–) 2–8 mm. 2n = 12.

Discussion

Varieties 5 (5 in the flora).

The diverse and intergrading forms here included in Chaenactis glabriuscula have been divided by P. Stockwell (1940) and subsequent workers into as many as four species and ten varieties. Chaenactis glabriuscula is known from the southern two-thirds of the Californian Floristic Province and adjacent desert edges. It has been reported in Massachusetts as a garden escape (variety unspecified); it is not expected to persist there outside cultivation.

Complete interfertility among the taxa recognized here as Chaenactis glabriuscula vars. glabriuscula, megacephala, and lanosa was demonstrated by P. Stockwell (1940). Intraspecific crosses involving C. glabriuscula var. orcuttiana were much less successful; C. glabriuscula var. heterocarpha was not tested. As noted by W. J. Hooker and G. A. W. Arnott ([1830–]1841) and D. W. Kyhos (1965), some forms of C. glabriuscula differ from C. stevioides or C. fremontii only in corolla color, which can be lost in older or poorly preserved specimens.

Selected References

None.

Key

1 Pappi of (5–)8 scales in 2, abruptly unequal series Chaenactis glabriuscula var. heterocarpha
1 Pappi of (1–)4(–5) scales in 1 series > 2
2 Proximal indument whitish, ± densely lanuginose; leaves ± basal (persistent) Chaenactis glabriuscula var. lanosa
2 Proximal indument grayish, ± arachnoid, or glabrescent; leaves basal (withering) and cauline > 3
3 Largest leaf blades succulent, mostly 2-pinnately lobed; stems ± horizontal Chaenactis glabriuscula var. orcuttiana
3 Largest leaf blades scarcely succulent, 1(–2)-pinnately lobed; stems ascending to erect > 4
4 Longest phyllaries 7–9 × 2–3 mm, outer glabrescent in fruit; pappi: lengths of longest scales ± 0.9(–1) times corollas Chaenactis glabriuscula var. megacephala
4 Longest phyllaries 5–7 × 1–2 mm, outer (at least medially) ± arachnoid-sericeous and, often, ± stipitate-glandular in fruit; pappi: lengths of longest scales 0.4–0.7 times corollas Chaenactis glabriuscula var. glabriuscula

"longest" is not a number.

... more about "Chaenactis glabriuscula"
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
paniculiform +, cymiform +  and corymbiform +
indeterminate +  and determinate +
James D. Morefield +
de Candolle in A. P. de Candolle and A. L. P. P. de Candolle +
decurrent +
compound +  and simple +
lobed;ovate;oblanceolate;ovate;oblanceolate;linear;deltate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
connate +  and distinct +
Yellow pincushion +
actinomorphic +
stipitate-glandular +  and hairy +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
fertile +  and bisexual +
fertile +  and bisexual +
Calif. +  and nw Mexico +
strigose +  and densely sericeous +
scabrellous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
campanulate +  and cylindric +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
less hemispheric;obconic or broadly cylindric +
lobed +  and plane +
not +  and succulent +
cauline +  and basal +
1 cm10 mm <br />0.01 m <br /> (?) +  and 10 cm100 mm <br />0.1 m <br /> (?) +
deltate +  and more or less lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
dentate to pinnatifid or palmatifid +  and entire +
glabrescent +, lanuginose +, arachnoid-sericeous +, villous +  and stipitate-glandular +
acute +  and blunt +
2-carpellate +
persistent +
coroniform +
20 cm200 mm <br />0.2 m <br /> (30 cm300 mm <br />0.3 m <br />) +
glabrescent +, lanuginose +, arachnoid-sericeous +, villous +  and stipitate-glandular +
1 cm10 mm <br />0.01 m <br /> (20 cm200 mm <br />0.2 m <br />) +
5 +  and 21 +
0.45 cm4.5 mm <br />0.0045 m <br /> (1 cm10 mm <br />0.01 m <br />) +
remote +  and more or less congested +
grayish +  and whitish +
arachnoid +  and densely lanuginose or glabrescent +
in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. +
bearing subulate enations +, hairy +  and bristly +
knobby;convex;more or less flat +
distinct +
8 +  and 4 +
unequal +
exalbuminous +
unequal +
5 (?) +  and 12 (?) +
usually erect +  and ascending +
1 +  and 5 +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Undefined (tribe Undefined) subtribe Bahiinae +  and Undefined (tribe Undefined) subtribe Palafoxiinae +
Chaenactis glabriuscula +
Chaenactis sect. Chaenactis +
species +
cylindric +
80 cm800 mm <br />0.8 m <br /> (300 cm3,000 mm <br />3 m <br />) +
1 cm10 mm <br />0.01 m <br /> (80 cm800 mm <br />0.8 m <br />) +
toothed +  and entire +
70 cm700 mm <br />0.7 m <br /> (200 cm2,000 mm <br />2 m <br />) +
5 cm50 mm <br />0.05 m <br /> (70 cm700 mm <br />0.7 m <br />) +