Carlquistia

B. G. Baldwin

Novon 9: 463. 1999.

Etymology: For Sherwin Carlquist, b. 1930, Californian botanist
Treatment appears in FNA Volume 21. Treatment on page 302. Mentioned on page 255, 298, 303.
 TaxonIllustrator 
FNA21 P48 Carlquistia muir.jpegCarlquistia muirii
Madia elegans
Madia sativa
Marjorie C. Leggitt
Barbara Alongi
Barbara Alongi

Perennials, 7–54 cm (rhizomatous, often matted). Stems (aerial) erect. Leaves mostly cauline; proximal opposite, distal alternate; sessile; blades lanceolate to linear, margins entire, faces hirsute to villous and glandular-pubescent. Heads discoid, borne singly or in loose, corymbiform arrays. Peduncular bracts: pit-glands, tack-glands, and/or spines 0 at tips. Involucres ± campanulate, 5–10+ mm diam. Phyllaries (modified paleae) (5–) 7–16 in 1 series, lanceolate to lance-linear, herbaceous, abaxially hirsute and glandular-pubescent. Receptacles flat, setulose, epaleate (except for bracts constituting “involucres”). Ray-florets 0. Disc-florets 7–29, bisexual, fertile; corollas yellow, tubes shorter than or about equaling funnelform throats, lobes 5, deltate (anthers yellow to brownish; styles glabrous proximal to branches). Cypselae ± terete (apices not beaked, faces scabrellous); pappi of 9–17 white to mauve or tawny, subulate, ± plumose scales (flattened bristles). x = 8.

Discussion

Species 1.

Carlquistia resembles another montane, perennial tarweed, Anisocarpus scabridus (2n = 14), and the two taxa were long treated as congeneric, in Raillardella or Raillardiopsis. Molecular phylogenetic evidence for a sister-group relationship between Carlquistia and Madia in the strict sense is consistent with x = 8 in both groups (B. G. Baldwin 1996). Carlquistia is treated as distinct from Madia on the basis of morphologic and ecologic disparity between the two groups.

Carlquistia is closely related to the Hawaiian silversword alliance (Argyroxiphium de Candolle, Dubautia Gaudichaud-Beaupré, and Wilkesia A. Gray); vigorous, largely sterile hybrids have been produced between C. muirii and Dubautia laevigata A. Gray, a large shrub of mesic forests in Kauai (B. G. Baldwin et al. 1991). Vigorous hybrids of low fertility also have been produced between C. muirii and the perennial California tarweeds Anisocarpus scabridus and Kyhosia bolanderi (D. W. Kyhos et al. 1990). Diploid pollen from a hybrid between Carlquistia and Kyhosia was successfully used in a cross with Dubautia scabra (de Candolle) D. D. Keck (G. D. Carr et al. 1996).

Lower Taxa

... more about "Carlquistia"
not tailed +, rounded +  and obtuse +
pale +  and dark +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
Bruce G. Baldwin +  and John L. Strother +
B. G. Baldwin +
decurrent +
compound +  and simple +
linear to linear-elliptic +
lanceolate;linear +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
hairy +  and glabrous +
fertile +  and bisexual +
fertile +  and bisexual +
For Sherwin Carlquist, b. 1930, Californian botanist +
glandular-pubescent +, hirsute +  and villous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
in loose , corymbiform arrays +  and singly +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
campanulate +
petiolate +  and sessile +
alternate +  and opposite +
deltate +
2-carpellate +
persistent +
glandular-pubescent +  and hirsute +
7 +  and 16 +
convex;conic +
baldwin1990a +
white;mauve or tawny +
9 +  and 17 +
plumose +  and subulate +
exalbuminous +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Carlquistia +
Asteraceae (tribe Heliantheae) subtribe Madiinae +
cylindric +
toothed +  and entire +
7 cm70 mm <br />0.07 m <br /> (54 cm540 mm <br />0.54 m <br />) +
shrub +  and subshrub +