Calycadenia fremontii

A. Gray in W. H. Emory

in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 100. 1859.

Endemic
Synonyms: Calycadenia ciliosa Greene Calycadenia elegans Greene Calycadenia pauciflora var. elegans Jepson
Treatment appears in FNA Volume 21. Treatment on page 275. Mentioned on page 271, 276.

Plants 10–100 cm; self-incompatible. Stems simple or branched (main axes usually obvious, branches relatively few, ± rigid), hispidulous to strigose (especially proximally, usually strigose with scattered, stouter hairs distally). Leaves mostly alternate, 2–8 cm (proximal), hispid to hispidulous and ± long-hairy (especially on margins and adaxial faces). Heads borne singly or in open, spiciform arrays (1–2+ per node). Peduncular bracts narrowly elliptic to oblanceolate, 2–10 mm (hispidulous, ± bristly and/or strongly pectinate-fimbriate) apices rounded or truncate, tack-glands 1–5+. Phyllaries 3–7 mm, abaxial faces ± hispidulous, ± bristly, shaggy long-hairy on distal margins, tack-glands (0–) 1–5+. Paleae 3–7 mm (receptacular cups ± campanulate, lengths often ± equaling diams.). Ray-florets (1–) 2–6; corollas white or cream to pinkish, or yellow (sometimes basally reddish), tubes 1–2 mm (± papillate, throats sometimes with dark red “eyes”), laminae 4.5–7 mm (central lobes ± equaling or narrower than laterals, mostly obovate, widest beyond middles, sinuses nearly equaling laminae). Disc-florets (4–) 6–21; corollas white to pinkish, or yellow, 4–5 mm. Ray cypselae 2.5–4 mm, smooth, glabrous or glabrate. Disc cypselae 2–4 mm (often ± angular), appressed-hairy; pappi of 8–14 long-aristate and short-blunt, fimbriate scales 1–3 mm (alternating). 2n = 12.


Phenology: Flowering spring–fall.
Habitat: Open, dry meadows, hillsides, gravelly outwashes, rocky barrens in chaparral associations
Elevation: 50–1400 m

Discussion

Calycadenia fremontii is variable and not easily divisible. It ranges from Siskiyou County through Mendocino, Lake, and Butte counties in California with one outlying population near Grants Pass in southern Oregon. Genetic change and chromosome repatterning have produced an extremely complex taxon that (as here circumscribed) includes populations with individuals to 20 cm and extensively branched with white corollas as well as populations with individuals to 100 cm and unbranched or with relatively few branches with yellow corollas. Some relatively small forms with white corollas are more closely related to tall forms with yellow corollas than they are to each other. At present, there is no known combination of characters that allows reasonably consistent delimitation of these or other elements within this assemblage of populations. The present circumscription combines three previously described species, including one traditionally considered an element of C. pauciflora. Additionally, C. fremontii intergrades with C. pauciflora (see discussion under 10. C. pauciflora).

Selected References

None.

Lower Taxa

None.
... more about "Calycadenia fremontii"
long-hairy +, shaggy +, bristly +  and hispidulous +
not tailed +, rounded +  and obtuse +
pale +  and dark +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
umbelliform +, spiciform +, paniculiform +, glomerulate +  and corymbiform +
indeterminate +  and determinate +
Robert L. Carr +  and Gerald D. Carr +
A. Gray in W. H. Emory +
decurrent +
compound +  and simple +
linear to linear-elliptic +
lobed;usually spatulate;oblanceolate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
white +  and pinkish or yellow +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
hairy +  and glabrous +
fusiform +, clavate +, obpyramidal +, terete +, compressed +  and obcompressed +
fertile +  and bisexual +
appressed-hairy +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
fertile +  and bisexual +
Calif. +  and Oreg. +
50–1400 m +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Open, dry meadows, hillsides, gravelly outwashes, rocky barrens in chaparral associations +
spiciform +  and corymbiform +
in open , spiciform arrays +  and singly +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
urceolate;obconic;urceolate;obconic;hemispheric;fusiform;globose;ellipsoid;cylindric;campanulate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
cauline +  and basal +
long-hairy +, hispid +  and hispidulous +
2 cm20 mm <br />0.02 m <br /> (?) +  and 8 cm80 mm <br />0.08 m <br /> (?) +
deltate +  and lanceovate +
dentate to pinnatifid or palmatifid +  and entire +
toothed +  and entire +
2-carpellate +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
persistent +
narrowly elliptic +  and oblanceolate +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Flowering spring–fall. +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
in W. H. Emory, Rep. U.S. Mex. Bound. +
glabrate +, glabrous +  and smooth +
obcompressed +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
convex;conic +
8 +  and 14 +
fimbriate +, short-blunt +  and long-aristate +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
exalbuminous +
branched +  and simple +
erect +  and spreading +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Calycadenia ciliosa +, Calycadenia elegans +  and Calycadenia pauciflora var. elegans +
Calycadenia fremontii +
Calycadenia +
species +
cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
toothed +  and entire +
5 cm50 mm <br />0.05 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
7 +, 6 +, 5 +  and 4 +