Fam. Pl. 2: 130. 1763.

Common names: Sea oxeye daisy seaside tansy or oxeye
Etymology: For Ole Borch (Olaus Borrichius), 1626–1690, Danish botanist
Treatment appears in FNA Volume 21. Treatment on page 129. Mentioned on page 65, 67.
FNA21 P21 Borrichia frutescens.jpegBorrichia arborescens
Pascalia glauca
Acmella repens
Marjorie C. Leggitt
Linda Ann Vorobik
Barbara Alongi

Perennials, subshrubs, or shrubs, to 150 cm (rhizomatous, forming clonal colonies, roots fibrous, sometimes adventitious). Stems decumbent to erect, branched ± throughout (sap sticky-resinous). Leaves cauline; opposite; ± petiolate or sessile; blades (usually 1-nerved, sometimes obscurely 3-nerved or 5-nerved) elliptic, linear, oblanceolate, obovate, or ovate (usually coriaceous or succulent), bases ± cuneate, margins entire or toothed (teeth often spine-tipped), faces glabrous or puberulent to villous and/or sericeous. Heads radiate, borne singly or (3–10) in cymiform arrays. Involucres globose or ovoid to hemispheric or broader, (5–) 8–13 (–18+) mm diam. Phyllaries persistent, 10–45 in 2–3 series (outer larger, elliptic to oblanceolate or ovate, apices acute, cuspidate, obtuse, rounded, or spine-tipped). Receptacles convex, paleate (paleae lanceolate to ovate, ± conduplicate, partially enclosing cypselae, apices often ± pungent). Ray-florets 7–30, pistillate, fertile; corollas yellow. Disc-florets 20–75, bisexual, fertile; corollas yellow, tubes shorter than funnelform throats, lobes 5, lance-triangular (anthers black with orange glands on connectives). Cypselae (gray to black) ± obcompressed, or obpyramidal and 3-angled or 4-angled (faces faintly finely reticulate); pappi persistent or tardily falling, (stramineous) coroniform or cupular (3–4-angled). x = 14.


United States, Mexico, West Indies


Species 3 (3 species, including 1 hybrid, in the flora).

Leaves of Borrichia species are usually heteroblastic: leaves on primary stems are usually ± petiolate (sometimes with spine-tipped teeth on margins of petioles or near bases of blades) and larger and relatively broader than the usually sessile leaves on secondary stems.


1 Leaves (at least mid cauline): margins entire or dentate to serrate (teeth remote, mostly toward apices), faces glabrous or sericeous; phyllaries (chartaceous in fruiting heads): apices acute, rounded, or obtuse; paleae: apices obtuse to acute Borrichia arborescens
1 Leaves (at least mid cauline): margins usually dentate to serrate, rarely entire, faces usually villous (longer hairs) and/or sericeous (shorter hairs) to glabrate, rarely glabrous; phyllaries: apices cuspidate or spine-tipped; paleae: apices cuspidate or spine-tipped > 2
2 Mid-cauline leaves (petioles or blade bases with spine-tipped teeth): margins dentate to serrate (usually only proximal 1/3–1/2, rarely most, of each margin); phyllaries (often reflexed, swollen in fruit): apices spine-tipped; paleae: apices spine-tipped Borrichia frutescens
2 Mid-cauline leaves: margins dentate or serrate (at least mid portions, often undulate as well); phyllaries: apices cuspidate; paleae: apices cuspidate Borrichia ×cubana

"broader" is not a number.

... more about "Borrichia"
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
John C. Semple +
Adanson +
bristlelike +
coroniform +
cuneate +
compound +  and simple +
ovate;obovate;ovate;obovate;oblanceolate;linear +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
Sea oxeye daisy +  and seaside tansy or oxeye +
hairy +  and glabrous +
4-angled +, 3-angled +, obpyramidal +  and obcompressed +
fertile +  and bisexual +
fertile +  and bisexual +
United States +, Mexico +  and West Indies +
For Ole Borch (Olaus Borrichius), 1626–1690, Danish botanist +
puberulent +  and villous and/or sericeous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
ovoid;hemispheric +
sessile +  and petiolate +
lance-triangular +
dentate to pinnatifid or palmatifid +  and entire +
toothed +  and entire +
2-carpellate +
persistent +
cupular +  and coroniform +
10 +  and 45 +
unequal +  and subequal +
bearing subulate enations +, hairy +  and bristly +
semple1977b +  and semple1978a +
exalbuminous +
decumbent +  and erect +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Undefined (tribe Undefined) subtribe Enceliinae +, Undefined (tribe Undefined) subtribe Engelmanniinae +, Undefined (tribe Undefined) subtribe Spilanthinae +, Undefined (tribe Undefined) subtribe Verbesininae +  and Undefined (tribe Undefined) subtribe Zinniinae +
Borrichia +
Asteraceae (tribe Heliantheae) subtribe Ecliptinae +
cylindric +
shorter to longer than campanulate +
toothed +  and entire +
shrub +  and subshrub +