Asteraceae (tribe Heliantheae) subtribe Varillinae

B. L. Turner & A. M. Powell in V. H. Heywood et al.

in V. H. Heywood et al., Biol. Chem. Compositae 2: 719. 1978.

Treatment appears in FNA Volume 21. Treatment on page 220. Mentioned on page 7, 251.

Subshrubs [shrubs], (10–) 20–30+ cm (sometimes rhizomatous). Leaves cauline; alternate [opposite]; ± sessile; blades linear to filiform (usually ± terete), margins entire, faces glabrous (not glanddotted). Heads discoid, borne singly [in corymbiform arrays]. Calyculi 0. Involucres obconic. Phyllaries persistent, [30–] 45–60 in [2–] 4–5 series (distinct, lance-linear, unequal, outer shorter, all resin-nerved). Receptacles conic, paleate (paleae falling, linear, flattish or weakly conduplicate, scarious, resin-nerved). Ray-florets 0. Disc-florets [25–] 60–150+, bisexual, fertile; corollas yellow, tubes shorter than or about equaling campanulate to cylindric throats, lobes 5, deltate; anther thecae pale; stigmatic papillae in 2 lines. Cypselae columnar to clavate, 8–15-ribbed, glabrous; pappi 0 [bristles].

Distribution

sw United States, Mexico

Discussion

Genus 1, species 2 (1 in the flora).

Varillinae may be allied with Clappiinae (H. Robinson 1981) or with Jaumeinae (P. O. Karis and O. Ryding 1994).

Selected References

None.

Lower Taxa

not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
spiciform +, racemiform +, paniculiform +  and corymbiform +
indeterminate +  and determinate +
Theodore M. Barkley† +, Luc Brouillet +  and John L. Strother +
B. L. Turner & A. M. Powell in V. H. Heywood et al. +
decurrent +
compound +  and simple +
linear;filiform +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
columnar +  and clavate +
fertile +  and bisexual +
fertile +  and bisexual +
sw United States +  and Mexico +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
deltate +
2-carpellate +
persistent +
45 +  and 60 +
in V. H. Heywood et al., Biol. Chem. Compositae +
styliferous +, neuter +  and pistillate +
bearing subulate enations +, hairy +  and bristly +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Asteraceae (tribe Heliantheae) subtribe Varillinae +
Asteraceae tribe Heliantheae +
subtribe +
campanulate +  and cylindric +
perennial +, biennial +  and annual +