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subshrubs, shrubs, trees, or vines [lianas], annual, biennial, or perennial, deciduous or evergreen, monoecious or dioecious; latex present or absent. Leaves alternate

serrate-dentate, or lobed. Inflorescences: flowers solitary or clustered in axillary or terminal glomerules or in short, cylindric spikes; bracts absent or 1–5, deciduous