View source for Madia ← Madia You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Madia |accepted_authority=Molina |publications={{Treatment/Publication |title=Sag. Stor. Nat. Chili, |place=136, 354. 1782 |year=1782 }} |basionyms= |synonyms= |hierarchy=Asteraceae;Asteraceae tribe Heliantheae;Asteraceae (tribe Heliantheae) subtribe Madiinae;Madia |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Asteraceae]]</div><div class="higher-taxon"><small>tribe</small>[[Asteraceae tribe Heliantheae]]</div><div class="higher-taxon"><small>subtribe</small>[[Asteraceae (tribe Heliantheae) subtribe Madiinae]]</div><div class="higher-taxon"><small>genus</small>[[Madia]]</div></div> |etymology=From native name in Chile |volume=Volume 21 |mention_page=page 255, 257, 295, 296, 298, 300, 302, 304 |treatment_page=page 303 }}<!-- --><span class="statement" id="st-d0_s0" data-properties="whole-organism some measurement;whole-organism duration"><b>Annuals,</b> 5–250 cm.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation"><b>Stems </b>erect.</span> <span class="statement" id="st-d0_s2" data-properties="leaf position;leaf position;leaf arrangement"><b>Leaves </b>mostly cauline (at flowering) proximal opposite (often in rosettes), distal alternate;</span> <span class="statement" id="st-d0_s3" data-properties="distal stem arrangement;distal stem architecture">sessile;</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade arrangement or shape or course;margin architecture or shape;margin shape;face pubescence;face pubescence">blades lanceolate or oblong-linear to linear, margins usually entire, sometimes toothed, faces hirsute to strigose, usually glandular-pubescent as well.</span> <span class="statement" id="st-d0_s5" data-properties="head architecture or arrangement;array architecture;array architecture;array architecture;array architecture"><b>Heads </b>usually radiate (sometimes discoid in M. glomerata), in corymbiform, paniculiform, racemiform, or spiciform arrays or in glomerules.</span> <span class="statement" id="st-d0_s6" data-properties="spine presence"><b>Peduncular </b>bracts: pit-glands, tack-glands, and/or spines 0.</span> <span class="statement" id="st-d0_s7" data-properties="involucre shape;involucre shape;involucre shape;involucre shape;involucre shape;involucre shape;involucre shape;involucre shape;involucre diameter"><b>Involucres </b>ellipsoid, depressed-globose, globose, obconic, ovoid, or urceolate, 1–10+ mm diam.</span> <span class="statement" id="st-d0_s8" data-properties="phyllary presence;phyllary quantity;series quantity"><b>Phyllaries </b>0 (then outer paleae functioning as phyllaries, sometimes in M. glomerata), or 1–22 in 1 series (lance-linear to lance-attenuate or oblanceolate, herbaceous, each mostly or wholly enveloping a subtended ray ovary, abaxially hirsute and, usually, glandular).</span> <span class="statement" id="st-d0_s9" data-properties="receptacle shape;receptacle pubescence;receptacle pubescence;receptacle architecture"><b>Receptacles </b>flat to convex, glabrous or setulose, paleate (paleae persistent or falling readily, in 1 series between rays and discs, ± connate or distinct, phyllary-like, more scarious).</span> <span class="statement" id="st-d0_s10" data-properties="ray-floret presence;ray-floret quantity;ray-floret architecture;ray-floret reproduction"><b>Ray-</b>florets 0 (sometimes in M. glomerata), or 1–22, pistillate, fertile;</span> <span class="statement" id="st-d0_s11" data-properties="corolla coloration">corollas yellowish (with maroon bases sometimes in M. elegans; purplish red sometimes in M. sativa).</span> <span class="statement" id="st-d0_s12" data-properties="disc-floret quantity;disc-floret reproduction;disc-floret reproduction;disc-floret architecture"><b>Disc-</b>florets 1–80+, bisexual and fertile or functionally staminate;</span> <span class="statement" id="st-d0_s13" data-properties="corolla coloration;corolla coloration;tube height or length or size;throat variability;lobe quantity;lobe shape">corollas usually yellow, sometimes purplish, tubes shorter than or about equaling funnelform throats, lobes 5, deltate (anthers ± dark purple or yellow to brownish; styles glabrous proximal to branches).</span> <span class="statement" id="st-d0_s14" data-properties="ray cypsela shape;ray cypsela shape;ray cypsela shape;ray cypsela shape;ray cypsela shape;ray cypsela shape"><b>Ray </b>cypselae compressed, ± 3-angled, or rarely terete, clavate (often arcuate, basal attachments central or offset, apices sometimes beaked, faces glabrous);</span> <span class="statement" id="st-d0_s15" data-properties="pappus presence">pappi 0.</span> <span class="statement" id="st-d0_s16" data-properties="disc cypsela shape;disc cypsela presence"><b>Disc </b>cypselae similar, sometimes obovoid (often ± straight, basal attachments central, apices not beaked), sometimes 0;</span> <span class="statement" id="st-d0_s17" data-properties="">pappi 0.</span> <span class="statement" id="st-d0_s18" data-properties="pappus presence;x chromosome quantity">x = 8.</span><!-- -->{{Treatment/Body |distribution=North America;South America;Pacific Islands (Hawaii);Pacific Islands (probably ) |discussion=<p>Species 10 (10 in the flora).</p><!-- --><p>Madia is more narrowly circumscribed here than in previous treatments by D. D. Keck (1959) and others. Molecular phylogenetic data have indicated that Madia in those earlier senses is not monophyletic (B. G. Baldwin 1996). As treated here, Madia comprises all members of Keck’s informal “section Madia” except M. minima (= Hemizonella) (Baldwin 1999b). Most species are reportedly either cross-incompatible or intersterile (J. Clausen 1951).</p> |tables= |references= }}<!-- --><div class="treatment-key"> ==Key== <div class="treatment-key-group"> {| class="wikitable fna-keytable" |-id=key-0-1 |1 |Ray laminae 6–19 mm; paleae readily falling; anthers yellow to brownish; ray cypselae beaked (beaks adaxially offset, curved) |[[Madia radiata|Madia radiata]] |-id=key-0-1 |1 |Ray laminae 0.7–20 mm (rays sometimes 0 in M. glomerata); paleae mostly persistent; anthers yellow to brownish or ± dark purple; ray cypselae sometimes beaked. |[[#key-0-2| > 2]] |-id=key-0-2 |2 |Plants self-incompatible (heads showy) or self-compatible (heads not showy); disc florets functionally staminate (cypselae 0, ovary walls remaining pallid, membranous) |[[#key-0-3| > 3]] |-id=key-0-2 |2 |Plants self-compatible (heads not showy); disc florets bisexual (forming cypselae, ovary walls becoming dark, rigid) |[[#key-0-4| > 4]] |-id=key-0-3 |3 |Ray laminae greenish yellow, 4–11 mm; anthers ± dark purple; ray cypselae ± 3-angled (abaxial sides broadly rounded, adaxial sides 2-faced, angles between those faces ca.70°), glossy |[[Madia citriodora|Madia citriodora]] |-id=key-0-3 |3 |Ray laminae bright yellow (sometimes with maroon bases), 4–20 mm; anthers yellow to brownish or ± dark purple; ray cypselae compressed (abaxial sides slightly rounded, adaxial sides 2-faced, angles between those faces 15–45°), dull or glossy |[[Madia elegans|Madia elegans]] |-id=key-0-4 |4 |Heads usually in glomerules, sometimes in corymbiform or paniculiform arrays; involucres narrowly ovoid or ellipsoid; ray florets 0 or 1–3 |[[Madia glomerata|Madia glomerata]] |-id=key-0-4 |4 |Heads in crowded or open, corymbiform, paniculiform, racemiform, or spiciform arrays; involucres depressed-globose, globose, obovoid, ovoid, or urceolate; ray florets (1–)3–13(–14) |[[#key-0-5| > 5]] |-id=key-0-5 |5 |Stems 1–30(–60) cm; phyllaries glandular-pubescent (glands golden yellow), apices ± erect, often sulcate; anthers yellow to brownish |[[#key-0-6| > 6]] |-id=key-0-5 |5 |Stems 10–200 cm; phyllaries glandular-pubescent (glands black, purple, or yellow), apices ± reflexed, flat; anthers ± dark purple |[[#key-0-7| > 7]] |-id=key-0-6 |6 |Heads in ± spiciform arrays; involucres globose or ovoid, 6–8 mm; disc florets5–15, corollas pubescent; ray cypselae sometimes purple-mottled, beakless |[[Madia subspicata|Madia subspicata]] |-id=key-0-6 |6 |Heads in racemiform or paniculiform arrays (peduncles filiform); involucres depressed-globose, 3–5 mm; disc florets 1(–2), corollas glabrous; ray cypselae black, beaked (beaks adaxially offset, curved) |[[Madia exigua|Madia exigua]] |-id=key-0-7 |7 |Ray cypselae black or purple, terete, glossy |[[Madia anomala|Madia anomala]] |-id=key-0-7 |7 |Ray cypselae black, purple, or mottled, compressed, dull or glossy |[[#key-0-8| > 8]] |-id=key-0-8 |8 |Stems glandular-pubescent throughout; heads in racemiform, paniculiform, or spiciform arrays; involucres 6–16 mm; mostly coastal |[[Madia sativa|Madia sativa]] |-id=key-0-8 |8 |Stems distally glandular-pubescent; heads in corymbiform, racemiform, or paniculiform arrays; involucres 5–10 mm; mostly away from immediate coast |[[#key-0-9| > 9]] |-id=key-0-9 |9 |Lateral branches often surpassing main stems (in large plants); ray laminae 6–8 mm |[[Madia citrigracilis|Madia citrigracilis]] |-id=key-0-9 |9 |Lateral branches seldom surpassing main stems; ray laminae 1.5–8 mm |[[Madia gracilis|Madia gracilis]] |} </div></div><!-- -->{{#Taxon: name=Madia |author=Bruce G. Baldwin;John L. Strother |authority=Molina |rank=genus |parent rank=subtribe |synonyms= |basionyms= |family=Asteraceae |illustrator=Barbara Alongi |illustration copyright=Flora of North America Association |distribution=North America;South America;Pacific Islands (Hawaii);Pacific Islands (probably ) |reference=None |publication title=Sag. Stor. Nat. Chili, |publication year=1782 |special status= |source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V19-20-21/V21_741.xml |tribe=Asteraceae tribe Heliantheae |subtribe=Asteraceae (tribe Heliantheae) subtribe Madiinae |genus=Madia |array architecture=spiciform;racemiform;paniculiform;corymbiform |blade arrangement or shape or course=oblong-linear to linear |blade shape=lanceolate |corolla coloration=purplish;yellow;yellowish |disc cypsela presence=absent |disc cypsela shape=obovoid |disc-floret architecture=staminate |disc-floret quantity=1;80 |disc-floret reproduction=fertile;bisexual |distal stem architecture=sessile |distal stem arrangement=alternate |face pubescence=glandular-pubescent;hirsute;strigose |head architecture or arrangement=radiate |involucre diameter=1mm;10mm |involucre shape=urceolate;ovoid;urceolate;ovoid;obconic;globose;depressed-globose;ellipsoid |leaf arrangement=opposite |leaf position=proximal;cauline |lobe quantity=5 |lobe shape=deltate |margin architecture or shape=entire |margin shape=toothed |pappus presence=absent;absent |phyllary presence=absent |phyllary quantity=1;22 |ray cypsela shape=clavate;terete;3-angled;terete;3-angled;compressed |ray-floret architecture=pistillate |ray-floret presence=absent |ray-floret quantity=1;22 |ray-floret reproduction=fertile |receptacle architecture=paleate |receptacle pubescence=setulose;glabrous |receptacle shape=flat;convex |series quantity=1 |spine presence=absent |stem orientation=erect |throat variability=equaling |tube height or length or size=shorter |whole-organism duration=annual |whole-organism some measurement=5cm;250cm |x chromosome quantity=8 }}<!-- -->[[Category:Treatment]][[Category:Asteraceae (tribe Heliantheae) subtribe Madiinae]] Templates used on this page: Template:Asteraceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Return to Madia.