Pennellia

Nieuwland

Amer. Midl. Naturalist 5: 224. 1918.

Etymology: For Francis Whittier Pennell, 1886–1952, American botanist
Basionym: Heterothrix (B. L. Robinson) Rydberg Bull. Torrey Bot. Club 34: 435. 1905,
Synonyms: Lamprophragma O. E. Schulz
Treatment appears in FNA Volume 7. Treatment on page 559. Mentioned on page 224, 226, 235, 243, 257, 561.
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 TaxonIllustrator 
FNA7 P69 Pennellia tricornuta.jpegPennellia tricornuta
Hesperis matronalis
Iberis sempervirens
Yevonn Wilson-Ramsey
Yevonn Wilson-Ramsey
Yevonn Wilson-Ramsey

Perennials; not scapose; pubescent or glabrous, trichomes simple, with stalked, forked or dendritic ones. Stems erect [ascending], often branched distally. Leaves basal and cauline; petiolate or sessile; basal (withered with age), rosulate, petiolate, blade margins entire, dentate to sinuate, or runcinate; cauline subsessile or sessile [shortly petiolate], blade margins entire [dentate to sinuate]. Racemes (several-flowered, sometimes secund, lax), considerably elongated in fruit. Fruiting pedicels erect, descending, ascending, or divaricate-ascending, slender, (glabrous). Flowers (usually actinomorphic, rarely zygomorphic, cupshaped); sepals erect, oblong to ovate, lateral pair slightly to strongly saccate basally, (usually pubescent, rarely glabrous); petals white (lavender or purple apically), spatulate to oblanceolate, (± longer than sepals), claw undifferentiated from blade, (apex obtuse); stamens slightly tetradynamous; filaments dilated basally; anthers ovate to oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens. Fruits siliques, sessile or shortly stipitate, linear, not torulose, straight or curved, terete or latiseptate; valves each with prominent or obscure midvein, glabrous or pubescent; replum rounded; septum complete; ovules 40–250 per ovary; style distinct or obsolete; stigma capitate. Seeds uniseriate or biseriate, plump or flattened, winged or not, oblong to ovate; seed-coat (reticulate-areolate), mucilaginous or not when wetted; cotyledons accumbent or incumbent. x = 8.

Distribution

sw United States, Mexico, Central America, South America

Discussion

Species 8 (3 in the flora).

Key

1 Fruits strongly latiseptate, 2-2.5 mm wide; seeds 2.4-2.8 mm, winged (broadly); septums opaque; ovules 40-50 per ovary; cotyledons accumbent. Pennellia tricornuta
1 Fruits terete or slightly latiseptate, 0.6-1.8 mm wide; seeds 0.7-1.3 mm, not winged; septums transparent; ovules 90-250 per ovary; cotyledons incumbent > 2
2 Racemes not secund; fruiting pedicels straight, erect or ascending; fruits (1.5-)2.2-5.8 cm, terete; ovules 90-140 per ovary; sepals 2.7-4 mm; petals 1.5-4(-5) mm. Pennellia micrantha
2 Racemes secund; fruiting pedicels arcuate, reflexed; fruits 5-10 cm, latiseptate; ovules 150-250 per ovary; sepals 4-9 mm; petals (4-)5-7(-12) mm. Pennellia longifolia

"elongated" is not a number."thick" is not a number.

... more about "Pennellia"
ovate +  and oblong +
Sara Fuentes-Soriano +  and Ihsan A. Al-Shehbaz +
Nieuwland +
Heterothrix +
appendaged +  and unappendaged +
not auriculate +
dentate +  and sinuate or runcinate +
much smaller +  and reduced +
urceolate +, campanulate +  and tubular +
branched +  and simple +
sessile +  and subsessile +
well-developed +
undifferentiated +
not +  and differentiated +
emarginate +  and entire +
incumbent +  and accumbent +
sw United States +, Mexico +, Central America +  and South America +
straight +  and curved +
For Francis Whittier Pennell, 1886–1952, American botanist +
not winged +  and unappendaged +
connate +  and distinct +
zygomorphic +  and actinomorphic +
2-valved +  and capsular +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
latiseptate +, stipitate +  and sessile +
curved +  and straight +
terete +  and not torulose +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
lobed;dentate;lobed;dentate +
obscure +  and prominent +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
rudimentary +
spatulate +  and oblanceolate +
trinucleate +  and 3(-11)-colpate +
Amer. Midl. Naturalist +
divaricate-ascending +, ascending +, descending +  and erect +
slender +
fuentes-soriano2004a +  and rollins1980a +
not +  and mucilaginous +
not +  and winged +
biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
oblong +  and ovate +
persistent +  and caducous +
distinct +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
procumbent +  and decumbent +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
Lamprophragma +
Pennellia +
Brassicaceae tribe Halimolobeae +
coiled +, 1-7-veined +  and veined +
pubescent +  and glabrous +
anastomosing +
glabrous +  and pubescent +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +