View source for Woodsia phillipsii

{{Treatment/ID
|accepted_name=Woodsia phillipsii
|accepted_authority=Windham
|publications={{Treatment/Publication
|title=Contr. Univ. Michigan Herb.
|place=19: 50. 1993
|year=1993
}}
|common_names=Phillips's cliff fern
|special_status={{Treatment/ID/Special_status
|code=F
|label=Illustrated
}}
|basionyms=
|synonyms=
|hierarchy=Dryopteridaceae;Woodsia;Woodsia phillipsii
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Woodsia]]</div><div class="higher-taxon"><small>species</small>[[Woodsia phillipsii]]</div></div>
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;petiole base quantity;petiole base duration"><b>Stems </b>compact to short-creeping, erect to horizontal, with few-to-many persistent petiole bases of unequal lengths;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale shape;central stripe coloration;margin coloration">scales mostly uniformly brown but at least some bicolored with dark central stripe and pale-brown margins, narrowly lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>5–35 × 1.5–6 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole coloration;petiole architecture;petiole fragility"><b>Petiole </b>light-brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade architecture or shape;blade architecture or function or pubescence;blade coating"><b>Blade </b>lanceolate, usually 2-pinnate proximally, sparsely to moderately glandular, never viscid;</span> <span class="statement" id="st-d0_s5" data-properties="stalk width;tip size">glandular-hairs with thin stalks and slightly expanded tips;</span> <span class="statement" id="st-d0_s6" data-properties="glandular-hair arrangement;scale shape">rachis with scattered glandular-hairs and hairlike scales.</span> <span class="statement" id="st-d0_s7" data-properties="pinna shape;pinna length or size;pinna shape;apex shape"><b>Pinnae </b>elongate-deltate to elliptic, longer than wide, often attenuate to a narrowly acute apex;</span> <span class="statement" id="st-d0_s8" data-properties="pair quantity;pinnule arrangement">largest pinnae with 7–18 pairs of widely spaced pinnules;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial and adaxial surface architecture or function or pubescence;abaxial and adaxial surface quantity">abaxial and adaxial surfaces somewhat glandular, lacking nonglandular hairs or scales.</span> <span class="statement" id="st-d0_s10" data-properties="pinnule architecture or shape;pinnule shape"><b>Pinnules </b>dentate, often shallowly lobed;</span> <span class="statement" id="st-d0_s11" data-properties="margin reflectance;margin size or width;margin architecture or pubescence or shape;gland quantity;projection architecture;projection coloration or reflectance;tooth due-to-presence;filament architecture">margins often lustrous adaxially, somewhat thickened, with occasional glands, appearing ciliate due to presence of multicellular translucent projections on teeth that often prolonged to form twisted filaments.</span> <span class="statement" id="st-d0_s12" data-properties="vein tip size;hydathode coloration;hydathode prominence"><b>Vein </b>tips usually enlarged to form whitish hydathodes visible adaxially.</span> <span class="statement" id="st-d0_s13" data-properties="indusium length;segment size or width;segment texture;cell architecture or shape;sporangium life cycle"><b>Indusia </b>of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, often greatly surpassing mature sporangia.</span> <span class="statement" id="st-d0_s14" data-properties="spore some measurement;2n chromosome quantity"><b>Spores </b>averaging 37–44 µm. <b>2n</b> = 76.</span><!--

-->{{Treatment/Body
|phenology=Sporulating summer–fall.
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
|elevation=1600–3200 m
|distribution=Ariz.;N.Mex.;Tex.;n Mexico
|discussion=<p>Woodsia phillipsii traditionally has been identified as W. mexicana. It differs from typical W. mexicana, however, in having completely filamentous indusial segments, multicellular (often filamentous) translucent projections on the pinnule margins, a greater number of pinnules per pinna, and a diploid chromosome number. Woodsia phillipsii is the only diploid species currently recognized in the W. mexicana complex, and it was probably involved in the hybrid origins of both W. mexicana and W. neomexicana. Some individuals of the latter species are difficult to distinguish from W. phillipsii (see comments under W. neomexicana), and the two taxa occasionally hybridize to produce sterile triploids of intermediate morphology. Woodsia phillipsii is also known to hybridize with W. plummerae (see comments under that species) and W. cochisensis.</p>
|tables=
|references=
}}<!--

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-->{{#Taxon:
name=Woodsia phillipsii
|authority=Windham
|rank=species
|parent rank=genus
|synonyms=
|basionyms=
|family=Dryopteridaceae
|phenology=Sporulating summer–fall.
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
|elevation=1600–3200 m
|distribution=Ariz.;N.Mex.;Tex.;n Mexico
|reference=None
|publication title=Contr. Univ. Michigan Herb.
|publication year=1993
|special status=Illustrated
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V2/V2_61.xml
|genus=Woodsia
|species=Woodsia phillipsii
|2n chromosome quantity=76
|abaxial and adaxial surface architecture or function or pubescence=glandular
|abaxial and adaxial surface quantity=lacking
|apex shape=acute
|blade architecture or function or pubescence=glandular
|blade architecture or shape=2-pinnate
|blade coating=viscid
|blade shape=lanceolate
|cell architecture or shape=isodiametric
|central stripe coloration=dark
|filament architecture=twisted
|gland quantity=occasional
|glandular-hair arrangement=scattered
|hydathode coloration=whitish
|hydathode prominence=visible
|indusium length=uniseriate
|leaf length=5cm;35cm
|leaf width=1.5cm;6cm
|margin architecture or pubescence or shape=ciliate
|margin coloration=pale-brown
|margin reflectance=lustrous
|margin size or width=thickened
|pair quantity=7;18
|petiole architecture=not articulate
|petiole base duration=persistent
|petiole base quantity=few-to-many
|petiole coloration=darker;straw-colored;light-brown
|petiole fragility=brittle
|pinna length or size=longer than wide
|pinna shape=attenuate;elongate-deltate;elliptic
|pinnule architecture or shape=dentate
|pinnule arrangement=spaced
|pinnule shape=lobed
|projection architecture=multicellular
|projection coloration or reflectance=translucent
|scale coloration=bicolored;brown
|scale shape=hairlike;lanceolate
|segment size or width=narrow
|segment texture=filamentous
|sporangium life cycle=mature
|spore some measurement=37um;44um
|stalk width=thin
|stem architecture or arrangement=compact
|stem orientation=erect;horizontal
|tip size=expanded
|tooth due-to-presence=prolonged
|vein tip size=enlarged
}}<!--

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