Wyethia

Nuttall

J. Acad. Nat. Sci. Philadelphia 7: 39, plate 5. 1834.

Common names: Mules-ears
Etymology: For Nathaniel Jarvis Wyeth, 1802–1856, early western American explorer
Treatment appears in FNA Volume 21. Treatment on page 100. Mentioned on page 65, 66.
 TaxonIllustrator 
FNA21 P14 Balsamorhiza rosea.jpegBalsamorhiza rosea
Scabrethia scabra subsp. scabra
Wyethia mollis
John Myers
John Myers
John Myers
FNA21 P15 Wyethia amplexicaulis.jpegWyethia amplexicaulis
Agnorhiza bolanderi
Verbesina virginica
John Myers
John Myers
Yevonn Wilson-Ramsey

Perennials, 15–60 (–100) cm (taproots relatively massive; caudices seldom branched). Stems erect (or bending erect from bases), branched mostly from bases. Leaves basal and cauline; alternate; usually petiolate (basal), sometimes sessile; blades (usually pinnately nerved, basal larger than cauline) ± deltate, elliptic-ovate, lanceolate, lance-elliptic, lance-linear, or oblong-ovate, bases truncate to cuneate, margins usually entire, rarely dentate to serrate (sometimes ciliate), faces glabrous or hairy (sometimes glanddotted or finely stipitate-glandular). Heads radiate, borne singly or 2–5 (–8+) in ± corymbiform to racemiform arrays. Involucres hemispheric to campanulate or turbinate, 12–60+ mm diam. Phyllaries persistent, 12–36 (–48) in 2–3+ series (subequal to unequal, outer sometimes foliaceous, much larger than inner). Receptacles flat to convex, paleate (paleae conduplicate, at least bases, papery). Ray-florets 5–25+, pistillate, fertile; corollas mostly yellow (cream to white in W. helianthoides). Disc-florets 35–150+, bisexual, fertile; corollas yellow, tubes much shorter than cylindric throats; lobes 5, ± deltate to lanceolate (style-branches stigmatic in 2 barely distinct lines, appendages ± filiform). Cypselae ± prismatic, weakly 3–4-angled (faces glabrous or hairy); pappi 0, or persistent, coroniform (usually lacerate) or of 1–4+ ovate to subulate, erose to lacerate (often basally connate) scales. x = 19.

Distribution

w North America

Discussion

Species 8 (8 in the flora).

Some species formerly included in Wyethia are here treated in Agnorhiza and Scabrethia. Balsamorhiza is closely related to Wyethia.

Key

1 Involucres 35–60+ mm diam.; outer phyllaries (30–)40–80+ mm (± foliaceous, much surpassing discs) > 2
1 Involucres 10–30(–40) mm diam.; outer phyllaries 15–30(–40) mm (not foliaceous, seldom surpassing discs) > 3
2 Leaf blades: faces glabrous or finely stipitate-glandular, sometimes sparsely pilosulous as well (usually shining); cypselae 10–13 mm Wyethia glabra
2 Leaf blades: faces densely tomentose to tomentulose (usually gland-dotted as well), glabrescent; cypselae 12–15 mm Wyethia helenioides
3 Leaf blades: faces tomentose to tomentulose (at least when young, glabrescent, usuallygland-dotted) Wyethia mollis
3 Leaf blades: faces glabrous, glabrate, hirsute, hirsutulous, hirtellous, pilose, pilosulous, scabrellous, or strigillose (not tomentose to tomentulose, often gland-dotted or stipitate-glandular) > 4
4 Leaf blades: faces glabrous or glabrate (gland-dotted); phyllary margins not ciliate; cypselae glabrous > 5
4 Leaf blades: faces usually hirsute, hirsutulous, hirtellous, pilose, pilosulous, scabrellous, or strigillose (sometimes vernicose, rarely finely stipitate-glandular or gland-dotted), sometimes glabrate; phyllary margins ± ciliate; cypselae strigillose or glabrous > 6
5 Basal leaves: blades lance-elliptic or oblong-lanceolate to lanceolate; rays 8–21(–25), laminae 25–60 mm; cypselae 8–9 mm Wyethia amplexicaulis
5 Basal leaves: blades narrowly oblong-lanceolate to lance-linear; rays 5–8+, lami-nae 18–30 mm; cypselae 6–8 mm Wyethia longicaulis
6 Basal leaves: blades lanceolate to lance-linear; cypselae 7–8 mm, ± strigillose Wyethia angustifolia
6 Basal leaves: blades elliptic-ovate, lanceolate, or lance-elliptic; cypselae 9–11 mm, strigillose or glabrous > 7
7 Phyllaries 36–48; rays 13–25, laminae (cream to white) 25–45 mm; cypselae strigillose distally and on margins Wyethia helianthoides
7 Phyllaries 16–34; rays 11–12, laminae (yellow) 25–50 mm; cypselaeglabrous Wyethia arizonica

"broader" is not a number.

... more about "Wyethia"
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
less corymbiform +  and racemiform +
indeterminate +  and determinate +
William A. Weber +
Nuttall +
bristlelike +
coroniform +
truncate +  and cuneate +
compound +  and simple +
oblong-ovate;lance-linear;oblong-ovate;lance-linear;lanceolate;elliptic-ovate;deltate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
Mules-ears +
hairy +  and glabrous +
3-4-angled +  and prismatic +
fertile +  and bisexual +
fertile +  and bisexual +
w North America +
For Nathaniel Jarvis Wyeth, 1802–1856, early western American explorer +
hairy +  and glabrous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
1.2 cm12 mm <br />0.012 m <br /> (6 cm60 mm <br />0.06 m <br />) +
hemispheric;campanulate or turbinate +
sessile +  and petiolate +
cauline +  and basal +
less deltate +  and lanceolate +
dentate +  and entire +
2-carpellate +
persistent +
of 1-4+ ovate to subulate , erose to lacerate scales +  and coroniform +
12 +  and 36 +
unequal +  and subequal +
J. Acad. Nat. Sci. Philadelphia +
bearing subulate enations +, hairy +  and bristly +
flat;convex +
weber1946a +
1 +  and 4 +
lacerate +, ovate +  and subulate +
exalbuminous +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Undefined (tribe Undefined) subtribe Enceliinae +, Undefined (tribe Undefined) subtribe Engelmanniinae +, Undefined (tribe Undefined) subtribe Spilanthinae +, Undefined (tribe Undefined) subtribe Verbesininae +  and Undefined (tribe Undefined) subtribe Zinniinae +
Wyethia +
Asteraceae (tribe Heliantheae) subtribe Ecliptinae +
cylindric +
cylindric +
shorter to longer than campanulate +
toothed +  and entire +
60 cm600 mm <br />0.6 m <br /> (100 cm1,000 mm <br />1 m <br />) +
15 cm150 mm <br />0.15 m <br /> (60 cm600 mm <br />0.6 m <br />) +
200 cm2,000 mm <br />2 m <br /> (400 cm4,000 mm <br />4 m <br />) +
shrub +  and subshrub +