Taraxacum ceratophorum

(Ledebour) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle

in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 7: 146. 1838.

Common names: Horned dandelion pissenlit tuberculé
Illustrated
Basionym: Leontodon ceratophorus Ledebour Icon. Pl. 1: 9, plate 34. 1829
Synonyms: Taraxacum ambigens var. fultius Fernald Taraxacum angulatum G. E. Haglund Taraxacum arctogenum Dahlstedt Taraxacum brachyceras Dahlstedt Taraxacum carthamopsis A. E. Porsild Taraxacum coverum R. Doll Taraxacum dumetorum Greene Taraxacum eriophorum Rydberg Taraxacum eurylepium Dahlstedt Taraxacum groenlandicum Dahlstedt Taraxacum hyperboreum Dahlstedt Taraxacum integratiforme R. Doll Taraxacum integratum G. E. Haglund Taraxacum lacerum Greene Taraxacum lateritium Dahlstedt Taraxacum longii Fernald Taraxacum mackenziense A. E. Porsild Taraxacum malteanum Dahlstedt ex G. E. Haglund Taraxacum maurolepium G. E. Haglund Taraxacum microcerum R. Doll Taraxacum ovinum Greene Taraxacum pellianum A. E. Porsild Taraxacum pseudonorvegicum Dahlstedt ex G. E. Haglund Taraxacum umbriniforme R. Doll Taraxacum umbrinum Dahlstedt ex G. E. Haglund
Treatment appears in FNA Volume 19. Treatment on page 247. Mentioned on page 241, 242, 248, 249.

Plants (1–) 6–50 cm; taproots branched. Stems 1–10+, ascending to erect, ± purplish (at least proximally), densely villous (young) becoming glabrescent, sparsely villous to glabrate or glabrous proximally, ± densely villous distally. Leaves ± 10, horizontal to patent, sometimes erect; sessile or petioles ± broadly winged (bases barely narrowed compared to blades); blades narrowly oblanceolate to linear-oblanceolate or linear-oblong (often ± runcinate), 4–30 × (0.4–) 0.5–5 cm, bases cuneate to attenuate, margins lobed ± deeply to lacerate, irregularly to regularly, often toothed, merely denticulate, or subentire, lobes retrorse, straight or antrorse, deltate to triangular, acute to acuminate, teeth 0–1 on lobes, often more or mostly in sinuses, apices obtuse to sometimes acute, sometimes mucronate, faces glabrous or glabrate to very sparsely villous. Calyculi of 12–16 (–20), appressed to spreading, pale, ovate to elliptic or lanceovate to lanceolate (sometimes thin) bractlets in 2–3 series, 5–12 × (0.9–) 1.5–5 mm, margins hyaline, white or purplish, scarious, apices caudate to acuminate, ± strongly horned, callous, or occasionally some (rarely all) hornless, tips obtuse to rounded, scarious, erose. Involucres dark green, sometimes ± glaucous, campanulate to ± hemispheric, (5–) 8–19 (–21) mm. Phyllaries (10–) 12–14 (–17) in 2 series, lanceolate to ovatelanceolate (inner), 1.5–4.5 mm wide, margins scarious or not (outer), inner broadly scarious in proximal 1/2, apices usually horned, occasionally hornless, horns sometimes exceeding apices, tips white to purplish, scarious, erose. Florets 40–85+; corollas yellow, drying cream to whitish (outer abaxially gray or purple-striped on drying), 10–22 × 1–2.8 mm. Cypselae olivaceous to olive brown, tan to olivaceous tan, brown to reddish-brown, grayish brown or straw-colored, bodies oblanceoloid to obovoid, 2.5–4 (–5) mm wide, cones conic or narrowly conic to broadly terete, 0.5–0.9 mm, beaks slender, 4.5–14 mm, ribs 5, large (bearing 10–15 narrower ones), faces proximally tuberculate or sometimes nearly smooth (usually with at least some tubercules) to muricate in distal 1/3–1/2, sometimes wholly muricate; pappi white to cream, 5–7.5 (–8) mm. 2n = 16, [24], 32, 40, 48.


Phenology: Flowering spring–summer.
Habitat: Wet to moist areas, calcareous or igneous rocks, gravel, sand, or clay, wet meadows, shores of streams, sandy or gravelly seashores, seepage slopes, early-melting snowbeds (south)
Elevation: 0–3000 m

Distribution

V19-323-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr., N.W.T., Nunavut, Ont., Que., Sask., Yukon, Alaska, Calif., Colo., Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo., Eurasia

Discussion

Taraxacum ceratophorum is the most widespread native dandelion in North America, ranging from the low Arctic and boreal zone to the western Cordilleras, in the montane and alpine zones.

This complex has been subdivided into many microspecies in North America, most of which appear unworthy of recognition. In the Quebec-Labrador Peninsula, Taraxacum ceratophorum grades continuously into what has been called T. hyperboreum. Inclusion of T. hyperboreum bridges the gap between typical T. ceratophorum and T. lacerum. Taraxacum lacerum stands out by its very lacerate leaves, but intermediates exist and it is impossible to draw a firm boundary. The lacerate Newfoundland form, T. longii, may be a spontaneous mutation within the range of T. ceratophorum. If T. lacerum were recognized, we would have to place T. longii within the former based on leaf morphology, though leaf orientation would be odd there (T. lacerum tends to have ascending leaves, and T. longii leaves that are flatter on the substrate). A more thorough morphometric and biosystematic study of this complex is warranted. Nonetheless, I have recognized two segregates (T. laurentianum, T. trigonolobum) that stand out from the continuum otherwise observed in the complex.

Selected References

None.

Lower Taxa

None.

"fine" is not a number.

... more about "Taraxacum ceratophorum"
48 +, 40 +, 32 +  and 16 +
hornless;horned;hornless;horned;caudate;acuminate +
ovate +  and lanceolate +
paniculiform +  and corymbiform +
indeterminate +  and determinate +
Luc Brouillet +
(Ledebour) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle +
cuneate +  and attenuate +
Leontodon ceratophorus +
slender +
0.45 cm4.5 mm <br />0.0045 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
compound +  and simple +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
4 cm40 mm <br />0.04 m <br /> (30 cm300 mm <br />0.3 m <br />) +
narrowly oblanceolate;linear-oblanceolate or linear-oblong +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
tuberculate +, rugose +, muricate +  and smooth +
oblanceoloid;obovoid +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
appressed +  and spreading +
ovate;elliptic or lanceovate +
filiform +
white +  and cream-colored or yellowish +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Horned dandelion +  and pissenlit tuberculé +
narrowly conic +  and broadly terete +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
zygomorphic +, actinomorphic +  and (3-)5-merous +
cream +  and whitish +
1 cm10 mm <br />0.01 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
olivaceous +  and olive brown tan +
tapered +, beaked +, flattened +, obcompressed +, compressed +, prismatic +, fusiform +, ellipsoid +, columnar +  and clavate +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Calif. +, Colo. +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +, Wyo. +  and Eurasia +
0–3000 m +
villous +, glabrate +  and glabrous +
muricate +, sometimes nearly smooth to muricate +  and tuberculate +
winged +  and beaked +
dispersed +
Wet to moist areas, calcareous or igneous rocks, gravel, sand, or clay, wet meadows, shores of streams, sandy or gravelly seashores, seepage slopes, early-melting snowbeds (south) +
indeterminate +
each +  and sessile +
1.9 cm19 mm <br />0.019 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (4 cm40 mm <br />0.04 m <br />) +
campanulate;more or less hemispheric +
0.8 cm8 mm <br />0.008 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
erect +, horizontal +  and patent +
subentire +, denticulate +, toothed +  and lacerate +
purplish +, white +  and hyaline +
deltate +  and triangular acute +
not +  and scarious +
2-carpellate +
0.75 cm7.5 mm <br />0.0075 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
white +  and cream +
persistent +
0.5 cm5 mm <br />0.005 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
Flowering spring–summer. +
closing +  and later +
distinct +  and coherent +
12 +  and 14 +
subequal +  and unequal +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. +
bearing subulate enations +, hairy +  and bristly +
flat;convex +
large +
exalbuminous +
Illustrated +
1 (?) +  and 10 (?) +
ascending +  and erect +
villous +, sparsely villous +  and glabrate or glabrous proximally +
1 +  and 10 +
scapiform +
appendaged +  and truncate +
papillate +
Taraxacum ambigens var. fultius +, Taraxacum angulatum +, Taraxacum arctogenum +, Taraxacum brachyceras +, Taraxacum carthamopsis +, Taraxacum coverum +, Taraxacum dumetorum +, Taraxacum eriophorum +, Taraxacum eurylepium +, Taraxacum groenlandicum +, Taraxacum hyperboreum +, Taraxacum integratiforme +, Taraxacum integratum +, Taraxacum lacerum +, Taraxacum lateritium +, Taraxacum longii +, Taraxacum mackenziense +, Taraxacum malteanum +, Taraxacum maurolepium +, Taraxacum microcerum +, Taraxacum ovinum +, Taraxacum pellianum +, Taraxacum pseudonorvegicum +, Taraxacum umbriniforme +  and Taraxacum umbrinum +
Taraxacum ceratophorum +
Taraxacum +
species +
white +  and purplish +
obtuse +  and rounded +
scarious +
with branched caudices +  and taprooted +
10 cm100 mm <br />0.1 m <br /> (30 cm300 mm <br />0.3 m <br />) +
30 cm300 mm <br />0.3 m <br /> (400 cm4,000 mm <br />4 m <br />) +
1 cm10 mm <br />0.01 m <br /> (6 cm60 mm <br />0.06 m <br />) +