Sidalcea oregana subsp. spicata

(Regel) C. L. Hitchcock

Perenn. Sp. Sidalcea, 64. 1957.

Common names: Spiked or spicate checkerbloom
EndemicIllustrated
Basionym: Callirhoe spicata Regel Gartenflora 21: 291, plate 737, figs. 3, 4. 1872
Synonyms: Sidalcea oregana var. spicata (Regel) Jepson S. spicata S. spicata var. tonsa M. Peck
Treatment appears in FNA Volume 6. Treatment on page 350. Mentioned on page 330, 334, 349, 351, 354.

Plants 0.3–0.8 (–0.9) m, with thick taproot, woody, branching crown. Stems branched or not, solid, base usually softly bristly-hirsute, hairs 1–2 mm, or stellate-hairy, sometimes glabrous. Leaves: stipules linear or lanceolate, 4–6 (–9) × 0.5–1 mm; petioles of basal leaves (5–) 10–20 (–30) cm, 3–4+ times as long as blade, reduced distally and distalmost leaves nearly sessile; blade reniform-orbiculate, cordate, unlobed and margins merely crenate or shallowly 5–7 (–9) -lobed, progressively more deeply lobed distally, 3–10 (–12) × 3–13 cm, lobes ternate or pinnately lobed, distalmost blades 3–5-lobed to base, lobes narrowly lanceolate, margins often entire, surfaces glabrous or sparsely appressed-hairy with simple hairs and/or minute, stellate hairs. Inflorescences spiciform, rachis apex usually acute in bud, usually dense, calyces overlapping, unbranched or branched, 20+-flowered, 10–15 cm, overlapping in flower and fruit (less often fruits spaced); primary peduncles usually equaling or shorter than mature inflorescence; bracts linear, undivided, 4–6 mm, usually longer than young flower buds and conspicuous, much longer than pedicels, shorter than to nearly equaling calyx. Pedicels 1–2 (–3) mm. Flowers bisexual or unisexual and pistillate, plants gynodioecious; calyx (3.5–) 5–6 (–7) mm, not accrescent, usually densely stellate-puberulent, sometimes bristly with hairs 0.5–1 (–2.5) mm; petals pink to rose-pink or magenta, pistillate 6–10 mm, bisexual 10–15 (–20) mm; staminal column 4–6 mm, hairy; anthers white to pinkish; stigmas 7–9. Schizocarps 4–7 mm diam.; mericarps 7–9, 2.5–3 mm, back moderately glandularpuberulent, usually smooth, margins rounded, sides sometimes lightly reticulate-veined, mucro 0.1–0.5 mm. Seeds 2 mm. 2n = 20, 40.


Phenology: Flowering (Jun–)Jul–Aug(–Sep).
Habitat: Moist, open meadows, streamsides, subalpine forests, yellow pine slopes
Elevation: 1100–3000 m

Discussion

Subspecies spicata has frequently been recognized as a distinct species but it clearly fits within the Sidalcea oregana complex. The extremes are readily recognizable, but the variation often makes for confusion with other subspecies, as well as with S. setosa. Similar species and subspecies have either been confused with it or included within it, including subspp. eximia, hydrophila, and valida, and even S. nelsoniana. Subspecies spicata is distinguished primarily by its elongated bud bracts, dense spicate inflorescences, essentially smooth mericarps, relatively small calyx, spreading-hirsute stem base, and lack of rhizomes. It is more restricted in range and elevation than subsp. oregana and occurs in the mountains from Mono County, California, north to Jefferson and Union counties, Oregon, and to Douglas and Washoe counties, Nevada.

Selected References

None.

Lower Taxa

None.

"elongating" is not a number.

white +  and pinkish +
Steven R. Hill +
(Regel) C. L. Hitchcock +
bristly-hirsute +
cordate +
Callirhoe spicata +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
unlobed +, cordate +  and reniform-orbiculate +
3cm;10cm +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
much longer +  and usually longer +
conspicuous +
divided +, undivided +  and 2-fid +
20+-flowered +, branched +  and unbranched +
overlapping +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
bristly +  and stellate-puberulent +
not accrescent +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
indehiscent +  and loculicidal +
Spiked or spicate checkerbloom +
3(-5)-lobed +  and unlobed +
Calif. +, Nev. +  and Oreg. +
1100–3000 m +
sparse;copious +
unisexual +  and bisexual +
Moist, open meadows, streamsides, subalpine forests, yellow pine slopes +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
glabrous +  and stellate-hairy +
stellate +  and forked +
minute +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
axillary +  and terminal +
tough-fibrous +
connate +  and distinct +
simple +, stipulate +, sessile +, subsessile +  and petiolate +
distichous +  and alternate +
10 cm100 mm <br />0.1 m <br /> (35 cm350 mm <br />0.35 m <br />) +
3-5 times as long as blades +
7 cm70 mm <br />0.07 m <br /> (10 cm100 mm <br />0.1 m <br />) +
cauline +  and basal +
lanceolate +  and lobed +
reticulate-veined +  and smooth +
10 cm100 mm <br />0.1 m <br /> (12 cm120 mm <br />0.12 m <br />) +
3cm +  and 10cm +
rounded;lobed;5-7(-9)-lobed;crenate +
3cm +  and 13cm +
not stellate-hairy +  and glabrous +
wedge--shaped +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
(3-)5-9-lobed +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
3-40-carpellate +
(1-)2-many +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
overlapping +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
adnate +  and distinct +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
20 cm200 mm <br />0.2 m <br /> (30 cm300 mm <br />0.3 m <br />) +
3-4+ times as long as blade +
10 cm100 mm <br />0.1 m <br /> (20 cm200 mm <br />0.2 m <br />) +
Flowering (Jun–)Jul–Aug(–Sep). +
Perenn. Sp. Sidalcea, +
elongated +
4mm +  and 7mm +
hairy +  and glabrous +
reniform +
Lobed (?) +  and Unlobed (?) +
persistent +
reticulate-veined +
Endemic +  and Illustrated +
sessile +  and subsessile +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
solid +, not +  and branched +
7 +  and 9 +
filiform +, linear +  and capitate +
1-2 times number of carpels +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
unbranched +  and branched +
appressed-hairy +  and glabrous +
Sidalcea oregana var. spicata +, S. spicata +  and S. spicata var. tonsa +
Sidalcea oregana subsp. spicata +
Sidalcea oregana +
subspecies +
not +  and glaucous +