Sidalcea malviflora subsp. patula

C. L. Hitchcock

Perenn. Sp. Sidalcea, 21. 1957.

Common names: Torch-flowered or spreading checkerbloom
Conservation concernEndemic
Treatment appears in FNA Volume 6. Treatment on page 344. Mentioned on page 341, 345, 356.

Plants (0.2–) 0.5–0.6 (–0.9) m, usually with elongated, freely-rooting, sometimes matted rhizomes 2–5 mm diam. Stems erect or ascending, base decumbent, freely rooting, softly bristly-hairy proximally, stellate-hairy distally, usually not densely so. Leaves mostly basal, some cauline; stipules sometimes purplish, lanceolate to ovate, 5–6 (–9) × 1.5–3 mm; petioles of basal leaves 7–30 cm, 5–8 times as long as blades, gradually reduced distally to 1–2 times as long as blades, distalmost petioles 1/2–1 times as long as blade; blade 3–12 cm wide, basal unlobed and margins crenate to shallowly 7–9-lobed and margins coarsely dentate-crenate, distal more deeply incised to digitate, distalmost divided nearly to base, 5–7-lobed, lobes dentate to laciniate, surfaces densely stellate-hairy abaxially, usually hirsute adaxially, hairs simple or forked. Inflorescences dense, spicate to capitate, usually unbranched or with reduced lateral branches, 10–20+-flowered, 2–6 cm, to 15–20 (–30) cm in fruit, fruits congested or spaced, not overlapping on elongated axis, flowers obviously overlapping, stiffly erect, usually 5+ open at same time, proximalmost 1–3 flowers usually in leaf-axils and separated from spike above, flowers in pistillate plants smaller and more separated; bracts lanceolate, 5–7 × 2 mm, shorter or longer than pedicels, distal undivided or 2-fid, proximal separated to base. Pedicels usually 2–3 (–10) mm. Flowers: calyx 8–13 mm, hairy, hairs small, stellate and with longer, usually forked bristles; petals bright pinkish to rose-pink, drying dark purple, pale-veined at least when dry, pistillate 9–11 (–15) mm, bisexual 15–25 mm; staminal column 5–7 mm; anthers white; stigmas (6 or) 7 or 8. Schizocarps 6–8 mm diam.; mericarps (6 or) 7 or 8, 3.5–4 mm, glandularpuberulent, prominently reticulate-veined, rugose, and pitted, mucro 0.3–0.5 (–1) mm. Seeds 2.5 mm. 2n = 40.


Phenology: Flowering May–Jul.
Habitat: Open coastal forests, grassy areas, sometimes serpentine
Elevation: 0–700 m

Discussion

Of conservation concern.

Subspecies patula is rare and is distinguished by its dense, relatively short, spiciform inflorescences that elongate in fruit, its usually rough indument, its tendency to retain basal leaves, and its relatively long, freely rooting rhizomes that sometimes form mats. Some collections show a transition with subsp. rostrata, which usually has unlobed stem leaves, a denser tomentum, and shorter rhizomes, and with Sidalcea virgata, which is less hairy, has shorter rootstocks, and is more open-flowered. Molecular data support the Malviflora clade including subsp. patula but not including S. virgata (K. Andreasen, pers. comm.). Subspecies patula is a candidate for threatened or endangered listing in Oregon; fewer than 20 populations of it are known. It is known from coastal Del Norte and Humboldt counties in California and Curry County in Oregon.

Selected References

None.

Lower Taxa

None.
Steven R. Hill +
C. L. Hitchcock +
elongated +
stellate-hairy +  and bristly-hairy +
cordate +
Sida malviflora +
dissected +, lobed +  and unlobed +
3cm;12cm +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
shorter or longer +
2-fid +, undivided +  and lanceolate +
2 cm20 mm <br />0.02 m <br /> (?) +
pistillate +  and pale-veined +
overlapping +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
bright pink +  and lavender or dark rose-purple +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
indehiscent +  and loculicidal +
Torch-flowered or spreading checkerbloom +
sparsely hairy;glabrate +
1/2-1 times as long as blade +
Calif. +  and Oreg. +
0–700 m +
sparse;copious +
separated +  and overlapping +
unisexual +  and bisexual +
not overlapping +, spaced +  and congested +
Open coastal forests, grassy areas, sometimes serpentine +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
10-20+-flowered +, with reduced lateral branches +, unbranched +, spicate +  and capitate +
20 cm200 mm <br />0.2 m <br /> (30 cm300 mm <br />0.3 m <br />) +
ascending +  and erect +
axillary +  and terminal +
15 cm150 mm <br />0.15 m <br /> (20 cm200 mm <br />0.2 m <br />) +
tough-fibrous +
connate +  and distinct +
simple +, stipulate +, sessile +, subsessile +  and petiolate +
distichous +  and alternate +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
1-2 times as long as blades +  and 5-8 times as long as blades +
7 cm70 mm <br />0.07 m <br /> (30 cm300 mm <br />0.3 m <br />) +
dentate +  and laciniate +
crenate-serrate +  and dentate +
distalmost +
5-7-lobed;divided;incised;dentate-crenate;crenate;shallowly 7-9-lobed +
reticulate-veined +
glabrous +, glandular +, stellate-puberulent +  and hirtellous +
wedge--shaped +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
5-9-lobed +  and unlobed +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
3-40-carpellate +
(1-)2-many +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
pistillate +  and pale-veined +
1.1 cm11 mm <br />0.011 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
adnate +  and distinct +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Flowering May–Jul. +
smaller +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
4 cm40 mm <br />0.04 m <br /> (15 cm150 mm <br />0.15 m <br />) +
lobed +, unlobed +, reniform +  and orbiculate +
4 cm40 mm <br />0.04 m <br /> (12 cm120 mm <br />0.12 m <br />) +
Perenn. Sp. Sidalcea, +
freely-rooting +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
elongated +
not cordlike +
6mm +  and 8mm +
hairy +  and glabrous +
reniform +
persistent +
reticulate-veined +
honeycomb-pitted +
Conservation concern +  and Endemic +
sessile +  and subsessile +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
solid +, branched +, unbranched +  and rooting +
scattered +  and clustered +
ascending;erect +
8 +  and 7 +
filiform +, linear +  and capitate +
1-2 times number of carpels +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
hairy +  and ciliate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
unbranched +  and branched +
hirsute +  and stellate-hairy +
Nuttallia malviflora +
Sidalcea malviflora subsp. patula +
Sidalcea malviflora +
subspecies +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
monoecious +, dioecious +  and hermaphroditic +