Ptychostomum warneum

(Röhling) J. R. Spence

Phytologia 87: 22. 2005.

Basionym: Mnium caespiticium var. warneum Röhling Deutschl. Fl. ed. 2, 3: 94. 1813
Synonyms: Bryum warneum
Treatment appears in FNA Volume 28. Treatment on page 174. Mentioned on page 167.

Plants in dense or open turfs, red, green, or yellow-green. Stems 0.5–2 (–3) cm, fertile stems comose, innovations evenly foliate. Leaves yellow-green to red, crowded, strongly contorted to shrunken when dry, ovatelanceolate, flat, 0.5–2 mm, often gradually enlarged toward stem apex; base usually green, not decurrent; margins revolute proximally, limbidium strong, in 2 or 3 rows; apex acuminate; costa short-excurrent, awn slender; proximal laminal cells rectangular, 3–4: 1; medial and distal cells 18–22 µm wide, 2–3: 1, walls thin. Specialized asexual reproduction absent. Sexual condition usually polyoicous, sometimes autoicous. Seta purple or red, 2–4 (–5) cm, stout, straight to somewhat flexuose. Capsule yellowbrown or brown, pyriform, symmetric, 2–4 mm, mouth yellow; hypophysis slender; operculum long-conic; peristome reduced; exostome teeth yellow or brown basally, pale-yellow to hyaline distally, lamellae straight, pores absent near base along mid line; endostome weakly adherent to exostome, basal membrane 1/2 exostome height, segments narrowly perforate, cilia absent or rudimentary. Spores with size variable in same capsule, (32–) 36–48 (–50) µm, pale green or yellow.


Phenology: Capsules mature Jul–Aug.
Habitat: Damp soil
Elevation: low elevations (0-50 m)

Distribution

V28 278-distribution-map.gif

Greenland, Que., n Eurasia (Himalayan Nepal)

Discussion

Ptychostomum warneum is an arctic-boreal species that is distinguished from the related P. calophyllum by its pyriform brown capsule with a slender neck, generally larger spores, and acuminate leaves. The stems are sparsely radiculose; the costal awns are smooth. The species is known from only one location in Quebec, in the Mingan Islands. Capsules are needed for identification.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

... more about "Ptychostomum warneum"
acuminate +
John R. Spence +
(Röhling) J. R. Spence +
denticulate +
slender +
reduced +
curved +  and straight +
not decurrent +
Mnium caespiticium var. warneum +
cucullate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
brown +  and yellowbrown +
nutant +, inclined +  and suberect +
long-exserted +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
not inflated +
not differentiated +
appendiculate +
rudimentary +
short-excurrent +
subpercurrent +
short-rhomboidal +
Greenland +, Que. +  and n Eurasia (Himalayan Nepal) +
low elevations (0-50 m) +
hyaline;pale-yellow +
hyaline +, brown +, orange +  and yellow +
pale-yellow +  and hyaline +
Damp soil +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
polyoicous +, monoicous +, dioicous +  and sexual +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
yellow-green +  and red +
erect;erect-spreading +
ovatelanceolate +
enlarged +  and shrunken +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
2-stratose +  and 1-stratose +
revolute entire +  and denticulate +
1 (?) +, 2 (?) +  and 3 (?) +
rhomboidal +  and hexagonal +
short-rectangular +  and quadrate +
long-conic +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature Jul–Aug. +
differentiated +
smooth;papillose +
perforate +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
red +  and purple +
flexuose +  and straight to somewhat +
elongate +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
yellow +  and pale green +
germinating +
variable +
branching +  and foliate +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Bryum warneum +
Ptychostomum warneum +
Ptychostomum subg. Ptychostomum +
species +
lanceolate +
distinct +
not +  and porose +
straight +
specialized +
autoicous +, polyoicous +, sexual +  and asexual +
sinuate +
incrassate +
yellow-green +, green +  and red +
large;small to very +