Ptychostomum pallens

(Swartz) J. R. Spence

Phytologia 87: 21. 2005.

Illustrated
Basionym: Bryum pallens Swartz Monthly Rev. 34: 538. 1801
Treatment appears in FNA Volume 28. Treatment on page 172. Mentioned on page 168, 173.

Plants in dense or open turfs, pale-pink, red, or rarely dull green. Stems 1–4 (–6) cm, fertile stems comose, innovations evenly foliate. Leaves pink, red, or rarely green, crowded, contorted to shrunken when dry, ovate to broadly ovatelanceolate, flat or weakly concave, 1–3.5 (–4) mm, often gradually enlarged toward stem apex; base pink or green, somewhat to strongly and narrowly decurrent; margins revolute proximally, plane distally, limbidium strong, in 2 or 3 rows; apex acuminate; costa percurrent to short-excurrent, awn stout; proximal laminal cells short-rectangular, 2–3 (–4):1; medial and distal cells 17–24 µm wide, 2–3: 1, walls thin, not porose. Specialized asexual reproduction rare, by leaf-axil gemmae, brown-red to green, finely papillose. Sexual condition dioicous. Seta redbrown, 2–4 (–6) cm, stout, straight to somewhat flexuose. Capsule yellowbrown, elongate-pyriform to clavate, somewhat curved, (3–) 4–5 mm, mouth yellowbrown; median exothecial cells short-rectangular to quadrate, 20–40 µm, 1–2: 1; operculum conic, apiculate; peristome usually well developed; exostome teeth yellow basally, hyaline distally, lamellae straight, pores absent near base along mid line; endostome not adherent to exostome, basal membrane 1/2 exostome height, segments with ovate perforations, cilia appendiculate to nodose, occasionally short. Spores 16–20 (–22) µm, yellow to green.


Phenology: Capsules mature Jun–Sep.
Habitat: Wet soil, along streams, wetlands, calcareous
Elevation: low to high elevations (0-3000 m)

Distribution

V28 274-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Maine, Mich., Minn., Mont., Nev., N.Mex., N.Y., Oreg., Utah, Wash., Wyo., s South America, Eurasia

Discussion

Ptychostomum pallens is a highly variable arctic-boreal to north-temperate species, related to P. cernuum, P. meesioides, and P. rutilans. The pale pink color is diagnostic but not always present. The species is distinguished from P. rutilans by its much thinner walled non-porose laminal cells and well-developed peristome; from P. meesiodes by its shorter, less arcuate capsule and short exothecial cells; and from P. cernuum by its dioicous sexual condition. The plants have red to red-brown leaf costae.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

... more about "Ptychostomum pallens"
acuminate +
John R. Spence +
(Swartz) J. R. Spence +
denticulate +
reduced +
green;pink +
curved +  and straight +
decurrent +
Bryum pallens +
cucullate +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
nutant +, inclined +  and suberect +
long-exserted +
elongate-pyriform +  and clavate +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
not inflated +
not differentiated +
appendiculate +
percurrent +  and short-excurrent +
subpercurrent +
short-rhomboidal +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Maine +, Mich. +, Minn. +, Mont. +, Nev. +, N.Mex. +, N.Y. +, Oreg. +, Utah +, Wash. +, Wyo. +, s South America +  and Eurasia +
low to high elevations (0-3000 m) +
hyaline;pale-yellow +
not adherent +
hyaline +, brown +, orange +  and yellow +
hyaline +  and yellow +
Wet soil, along streams, wetlands, calcareous +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
brown-red +  and green +
polyoicous +, monoicous +, dioicous +  and sexual +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
green +, red +  and pink +
erect;erect-spreading +
ovate +  and broadly ovatelanceolate flat or weakly concave +
enlarged +  and shrunken +
0.1 cm1 mm <br />0.001 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
2-stratose +  and 1-stratose +
1 (?) +, 2 (?) +  and 3 (?) +
rhomboidal +  and hexagonal +
short-rectangular +  and quadrate +
yellowbrown +
short-rectangular +  and quadrate +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature Jun–Sep. +
differentiated +
smooth;papillose +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
flexuose +  and straight to somewhat +
elongate +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Illustrated +
yellow +  and green +
germinating +
variable +
branching +  and foliate +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Ptychostomum pallens +
Ptychostomum subg. Ptychostomum +
species +
lanceolate +
distinct +
straight +
specialized +
dioicous +  and sexual +
sinuate +
incrassate +
green +, dull +, red +  and pale-pink +
large;small to very +