Polypodium californicum

Kaulfuss

Enum. Filic. 102. 1824.

Common names: California polypody
Illustrated
Synonyms: Polypodium californicum var. kaulfussii D. C. Eaton Polypodium vulgare subsp. californicum (Kaulfuss) Hultén Polypodium vulgare var. kaulfussii (D. C. Eaton) Fernald
Treatment appears in FNA Volume 2.

Stems dull or inconspicuously glaucous, moderately stout, to 10 mm diam., acrid or bland-tasting; scales uniformly brown or slightly darker near point of attachment, lanceolate-ovate, symmetric, margins entire to erose. Leaves to 70 cm. Petiole usually slender, to 3 mm diam. Blade deltate to lanceolate-ovate, pinnatifid, usually widest near base, to 20 cm wide, leathery to herbaceous; rachis sparsely scaly to glabrescent abaxially, puberulent adaxially; scales deltate to ovate, usually more than 10 cells wide. Segments linear-lanceolate to oblong, usually less than 15 mm wide; margins serrate; apex widely obtuse to rarely attenuate; midrib puberulent adaxially. Venation weakly to conspicuously anastomosing, most segments containing several areoles, often forming 1 row. Sori midway between margin and midrib or slightly closer to midrib, usually less than 3 mm diam., oval when immature. Sporangiasters absent. Spores less than 58 µm, verrucose, with surface projections to 3 µm. 2n = 74.


Phenology: Sporulating early winter–spring.
Habitat: Cliffs and soil on rocky slopes, on a variety of substrates but usually igneous
Elevation: 0–1500 m.

Distribution

V2 193-distribution-map.gif

Calif., Mexico in Baja California

Discussion

R. M. Lloyd and F. A. Lang (1964) recognized two cytotypes within Polypodium californicum. The tetraploid has proved to be an allopolyploid involving P. californicum and P. glycyrrhiza and is treated here as a separate species, P. calirhiza, following S. A. Whitmore and A. R. Smith (1991). Polypodium californicum can be confused with P. calirhiza, but it usually can be distinguished by blade shape, venation, spore size, and geographic distribution. D. S. Barrington et al. (1986) reported that spores of northern populations of P. californicum can be as large as those of P. calirhiza, but the former species has veins forming more areoles per segment than does the latter.

Selected References

None.

Lower Taxa

None.

"widest" is not a number.

... more about "Polypodium californicum"
widely obtuse +  and rarely attenuate +
Christopher H. Haufler +, Michael D. Windham +, Frank A. Lang +  and S. A. Whitmore +
Kaulfuss +
0cm;20cm +
pinnate +  and pinnatisect +
scaly +  and not glaucous +
pinnatifid +, deltate +  and lanceolate-ovate +
leathery +  and herbaceous +
California polypody +
Calif. +  and Mexico in Baja California +
0–1500 m. +
elliptic +  and cordate +
Cliffs and soil on rocky slopes, on a variety of substrates but usually igneous +
monomorphic +
closely spaced;distant +
narrowed +
0 cm0 mm <br />0 m <br /> (90 cm900 mm <br />0.9 m <br />) +
entire +  and denticulate +
winged +, grooved +  and articulate +
straw-colored +
terete +  and flattened +
Sporulating early winter–spring. +
Enum. Filic. +
not adaxially +  and grooved +
puberulent;sparsely scaly;glabrescent abaxially +
clathrate;not peltate;not clathrate;strongly clathrate +
darker;brown +
deltate +  and ovate +
linear-lanceolate +  and oblong +
0 cm0 mm <br />0 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0 cm0 mm <br />0 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
discrete +
Illustrated +
of modified sporangia +
yellowish +  and transparent +
pruinose +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
few;numerous +
bland-tasting +  and acrid +
verrucose +  and tuberculate +
Polypodium californicum var. kaulfussii +, Polypodium vulgare subsp. californicum +  and Polypodium vulgare var. kaulfussii +
Polypodium californicum +
Polypodium +
species +
anastomosing +
epiphytic +  and terrestrial +
pendent +, arching +  and erect +