Polygonum polygaloides

Meisner in A. P. de Candolle and A. L. P. P. de Candolle

in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 101. 1856.

Common names: Polygala knotweed
IllustratedEndemic
Treatment appears in FNA Volume 5. Treatment on page 565. Mentioned on page 561, 562.

Herbs. Stems erect, green, usually divaricately branched, rarely simple, ± wiry, (2–) 6–20 (–25) cm, glabrous. Leaves uniformly distributed, articulated to ocreae, basal leaves often caducous, distal leaves abruptly reduced to bracts; ocrea 4–8 mm, glabrous, proximal part cylindric, distal part silvery, with inconspicuous veins, lacerate; petiole absent; blade 3-veined, lateral-veins sometimes inconspicuous, without pleats, narrowly linear, 10–40 × 1–2.5 mm, margins ± revolute, smooth, apex acute or mucronate. Inflorescences axillary and terminal, spikelike, subglobose to cylindric; cymes in most axils or crowded distally, 1–3-flowered. Pedicels enclosed in ocreae, erect, 0.1–2 mm, sometimes absent. Flowers mostly closed; perianth 1.5–3 mm; tube 19–40% of perianth length; tepals overlapping, uniformly white, pink, or red, petaloid, oblong-lanceolate, ± navicular, apex acute to acuminate; midveins usually unbranched or with 2 lateral branches proximally, moderately to strongly thickened, tepals appearing ± keeled; stamens 3–8. Achenes enclosed in perianth, light yellow, light-brown, or greenish brown to dark-brown, ovate to lanceolate, 1.3–2.5 mm, faces subequal, shiny or dull, smooth or reticulate with longitudinal ridges.

Distribution

V5 1154-distribution-map.gif

Alta., B.C., Sask., Ariz., Calif., Colo., Idaho, Mont., N.Mex., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Subspecies 4 (4 in the flora).

J. C. Hickman’s (1993c) treatment of the Polygonum polygaloides complex is provisionally accepted here. Most of the intermediate specimens occur between subspp. confertiflorum, esotericum, and kelloggii. Alternatively, P. polygaloides could be recognized in the narrow sense and the three other taxa could be treated as subspecies of a separate P. kelloggii, the earliest available binomial.

Selected References

None.

Key

1 Margins of bracts green, if white then scarious border less than 0.2 mm wide > 2
1 Margins of bracts white, scarious border (0.2-)0.25-1 mm wide > 3
2 Achenes lanceolate, 2-2.5 mm; bracts elliptic-lanceolate, appressed, rigid; stamens 5-8 Polygonum polygaloides subsp. esotericum
2 Achenes ovate, 1.3-1.7 mm; bracts linear to linear-lanceolate, ± patent, soft; stamens 3 Polygonum polygaloides subsp. kelloggii
3 Inflorescences narrowly cylindric, continuous from bases of branches or stems; achenes 2-2.5 mm, lanceolate Polygonum polygaloides subsp. esotericum
3 Inflorescences round to ovate or cylindric, confined to tips of branches, rarely continuous from bases of branches or stems; achenes 1.3-2.1 mm, ovate-lanceolate to ovate > 4
4 Inflorescences round to ovate; scarious borders of bracts 0.5-1 mm wide; stamens8 Polygonum polygaloides subsp. polygaloides
4 Inflorescences ovate to cylindric; scarious borders of bracts 0.25-0.4 mm wide; stamens 3 Polygonum polygaloides subsp. confertiflorum

"/2" is not declared as a valid unit of measurement for this property.

... more about "Polygonum polygaloides"
reticulate +, tubercled +  and smooth +
greenish brown;dark-brown +
exserted +  and included +
dull +  and shiny +
ovate +  and lanceolate +
pink +  and purple +
elliptic +  and oblong +
acute;acuminate +
Mihai Costea +, François J. Tardif +  and Harold R. Hinds† +
Meisner in A. P. de Candolle and A. L. P. P. de Candolle +
not stipelike +
narrowly linear;subround +
coriaceous +
foliaceous +  and perfoliate +
scale-like +
just below the soil surface or and above the soil surface +  and spreading +
Polygala knotweed +
1-3-flowered +
Alta. +, B.C. +, Sask. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, N.Mex. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
reticulate +  and smooth +
dull +  and shiny +
subequal +
adnate +, free +  and distinct +
semi-open +  and open +
disarticulating +  and solid +
decumbent to erect +  and prostrate +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
compound +, branched +  and simple +
subglobose +  and cylindric +
nonmembranous +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
articulated +
distributed +
deciduous +
not rugulose +
papillose-denticulate +
with 2 lateral branches +  and unbranched +
swollen +
silvery +, white +  and hyaline +
disintegrating +
deciduous +  and persistent +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
coriaceous +  and membranous +
somewhat larger +
anatropous +, , +  and orthotropous +
accrescent +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
spreading;erect +
greenish white;pink +
campanulate +  and urceolate +
nonaccrescent +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
fleshy +, indurate +  and membranous +
not +  and articulated +
(2-)3(-4)-carpellate +
in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. +
papillose-scabridulous +  and smooth +
slender;stout +
woody +  and fibrous +
Illustrated +  and Endemic +
3 +  and 8 +
wiry +, simple +  and branched +
20 cm200 mm <br />0.2 m <br /> (25 cm250 mm <br />0.25 m <br />) +
zigzagged +  and straight +
papillous-scabridulous +
4-gonous +
6 cm60 mm <br />0.06 m <br /> (20 cm200 mm <br />0.2 m <br />) +
2 +  and 3 +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
connate +  and distinct +
spreading;erect +
Undefined sect. Duravia +  and Polygonum sect. Monticola +
Polygonum polygaloides +
Polygonum sect. Duravia +
species +
overlapping +
red +, pink +  and white +
connate +  and distinct +
keeled +, navicular +  and oblong-lanceolate +
coriaceous +
persistent +
membranous +
pubescent +  and glabrous +
inconspicuous +
heterophyllous +  and homophyllous +