Penstemon barbatus

(Cavanilles) Roth

Catal. Bot. 3: 49. 1806.

Common names: Beard-lip beardtongue
Illustrated
Basionym: Chelone barbata Cavanilles Icon. 3: 22, plate 242. 1795
Treatment appears in FNA Volume 17. Treatment on page 146. Mentioned on page 145, 147.

Stems erect, sometimes ascending, 30–100 cm, glabrous or puberulent proximally, usually slightly glaucous. Leaves glabrous or sparsely to densely puberulent, not glaucous; basal and proximal cauline 30–80 (–140) × (6–) 12–30 (–35) mm, blade oblanceolate, base tapered, margins entire, apex obtuse, acute, or acuminate; cauline 3–7 (–14) pairs, sessile, 28–142 × 1–8 (–15) mm, blade lanceolate to linear, base tapered, apex acuminate, sometimes acute. Thyrses ± secund, sometimes cylindric, 11–69 cm, axis glabrous, verticillasters 6–14 (–19), cymes (1 or) 2–4-flowered; proximal bracts linear, 15–70 (–126) × 1–10 mm; peduncles and pedicels spreading to ascending, glabrous. Flowers: calyx lobes ovate to lanceolate, (3–) 4–6 (–9) × 1.9–2.8 (–3.2) mm, margins entire or erose, glabrous or obscurely glandular-pubescent; corolla red or orangish red to crimson or scarlet, with reddish violet nectar guides, bilaterally symmetric, strongly bilabiate, tubular-funnelform, 26–32 (–36) mm, glabrous or sparsely white-lanate or yellow-lanate internally abaxially, tube 5–8 mm, throat 5–6 mm diam., not constricted at orifice, slightly 2-ridged abaxially, abaxial lobes strongly reflexed, adaxial lobes projecting; stamens exserted (but usually concealed beneath projecting adaxial lip), pollen-sacs opposite, proximal 1/4–1/3 indehiscent, 1.5–2 (–2.2) mm, sides glabrous or sparsely to moderately lanate-villous, hairs white, to 1.5 mm, sutures papillate or denticulate, teeth to 0.1 mm; staminode 15–17 mm, included, tip straight, glabrous; style 22–27 mm. Capsules 10–14 × 6–8 mm.

Distribution

Ariz., Calif., Colo., N.Mex., Tex., Utah, n Mexico

Discussion

Varieties 3 (3 in the flora).

Penstemon barbatus is among the more widespread and conspicuous penstemons in the desert Southwest and Southern Rocky mountains. Putative hybrids reported between P. barbatus and P. comarrhenus, P. glaber, P. strictus, and P. virgatus were summarized by F. S. Crosswhite (1965); putative hybrids involving P. pseudoputus and P. putus also are suspected among some herbarium specimens. Penstemon barbatus is widely cultivated for its showy flowers; cultivars with yellow corollas also exist. Penstemon barbatus was reported for Suffolk County, Massachusetts (D. W. Magee and H. E. Ahles 2007); that report, or that those plants persisted outside of cultivation, has not been confirmed.

Various groups of Apache, Navajo, and Western Keres of Arizona and New Mexico used Penstemon barbatus for drugs, ceremonies, and decorations (D. E. Moerman 1998).

Selected References

None.

Key

1 Pollen sac sides sparsely to moderately lanate-villous. Penstemon barbatus var. trichander
1 Pollen sac sides glabrous. > 2
2 Corollas yellow-lanate internally abaxially. Penstemon barbatus var. barbatus
2 Corollas glabrous or sparsely white-lanate internally abaxially. Penstemon barbatus var. torreyi

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon barbatus"
acute +, acuminate +  and obtuse +
Craig C. Freeman +
(Cavanilles) Roth +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered;tapered +
Chelone barbata +
lanceolate;linear +
not +  and leathery +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
ovate;lanceolate +
1.9mm;2.8mm +
short-tubular +  and short-campanulate +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2.8 cm28 mm <br />0.028 m <br /> (14.2 cm142 mm <br />0.142 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
short-petiolate +  and sessile +
Beard-lip beardtongue +
3.2 cm32 mm <br />0.032 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
orangish red +  and crimson or scarlet +
yellow-lanate +, white-lanate +  and glabrous +
tubular-funnelform +
ventricose-ampliate +  and ampliate +
2.6 cm26 mm <br />0.026 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
2-4-flowered +
scale-like +  and not needlelike +
Ariz. +, Calif. +, Colo. +, N.Mex. +, Tex. +, Utah +  and n Mexico +
drupe-like +
curved +  and straight +
1 +  and 4 +
0 cm0 mm <br />0 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
not glaucous +, glabrous or +  and sparsely densely puberulent +
not +  and leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
glandular-pubescent +  and glabrous +
toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
spreading +  and ascending +
spreading;ascending +
5 +  and 4 +
0.2 cm2 mm <br />0.002 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
navicular-sigmoid +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1/4 +  and 1/3 +
7 cm70 mm <br />0.07 m <br /> (12.6 cm126 mm <br />0.126 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (7 cm70 mm <br />0.07 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Catal. Bot. +
minute +
brown +  and dark-brown +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.42 cm4.2 mm <br />0.0042 m <br />) +
glabrous or +  and sparsely moderately lanate-villous +
Illustrated +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (?) +  and 1.7 cm17 mm <br />0.017 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
glaucous +, puberulent +  and glabrous +
30 cm300 mm <br />0.3 m <br /> (100 cm1,000 mm <br />1 m <br />) +
2.2 cm22 mm <br />0.022 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
papillate +
denticulate +
Penstemon barbatus +
Penstemon sect. Elmigera +
species +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
2-ridged +
not constricted +
cylindric +
11 cm110 mm <br />0.11 m <br /> (69 cm690 mm <br />0.69 m <br />) +
straight +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
gibbous +  and not spurred +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
cauline +  and proximal +