Geum macrophyllum var. perincisum

(Rydberg) Raup

Rhodora 33: 176. 1931.

Endemic
Basionym: Geum perincisum Rydberg in N. L. Britton et al., N. Amer. Fl. 22: 405. 1913
Synonyms: G. macrophyllum subsp. perincisum (Rydberg) Hultén G. macrophyllum var. rydbergii Farwell G. oregonense G. perincisum var. intermedium B. Boivin
Treatment appears in FNA Volume 9. Treatment on page 66. Mentioned on page 65, 67, 68.

Cauline leaves: proximal pinnate or lyrate-pinnate, distal 3-foliolate or simple and 3-lobed (divided almost to base), lobes oblanceolate to obovate. Pedicels densely puberulent, sometimes also with scattered longer hairs, stipitate-glandular. 2n = 42.


Phenology: Flowering late spring–summer.
Habitat: Wet meadows, stream banks, thickets, muskegs, clearings, forest edges, roadsides
Elevation: 100–3100 m

Distribution

V9 97-distribution-map.jpg

Alta., B.C., Man., N.W.T., Nunavut, Ont., Que., Sask., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Mich., Minn., Mont., Nev., N.Mex., N.Dak., Oreg., S.Dak., Utah, Wash., Wis., Wyo.

Discussion

Most specimens of Geum macrophyllum with glands on the pedicels also have the more deeply dissected leaves typical of var. perincisum; exceptions occur. Plants collected in the San Bernardino Mountains and the southern Sierra Nevada of California are so densely glandular as to fit this variety, yet their leaves are like those of var. macrophyllum. More troublesome are specimens from the Upper Peninsula of Michigan with sparsely glandular pedicels and leaves of var. macrophyllum. These plants are likely hybrids between the two varieties as their ranges overlap at the western end of the Great Lakes region. Occasional intermediate specimens were also seen from northern California to British Columbia where the ranges of the two varieties meet. Specimens of var. perincisum from around James Bay have deeply dissected leaves, but some have glandless pedicels and occasionally the styles are completely glabrous.

Variety perincisum hybridizes with Geum rivale (= G. ×pervale B. Boivin); see discussion under 8. G. rivale.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."elongating" is not a number. "elongating" is not a number.

stipitate-glandular +
hooked;rounded +
Joseph R. Rohrer +
(Rydberg) Raup +
compound +  and simple +
opposite +  and alternate +
10 cm100 mm <br />0.1 m <br /> (45 cm450 mm <br />0.45 m <br />) +
Geum perincisum +
simple +, 3-foliolate +  and pinnate +
lyrate-pinnate;lyrate-pinnate +
3-lobed +
coriaceous +
free +  and distinct +
2 cm20 mm <br />0.02 m <br /> (12 cm120 mm <br />0.12 m <br />) +
simple +  and 3-foliolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Alta. +, B.C. +, Man. +, N.W.T. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mich. +, Minn. +, Mont. +, Nev. +, N.Mex. +, N.Dak. +, Oreg. +, S.Dak. +, Utah +, Wash. +, Wis. +  and Wyo. +
not +  and aggregated +
100–3100 m +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (4.6 cm46 mm <br />0.046 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
tapered +, ovoid +  and fusiform +
0.2 cm2 mm <br />0.002 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Wet meadows, stream banks, thickets, muskegs, clearings, forest edges, roadsides +
saucer-shaped +  and cupshaped +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
crassinucellate +
odd-pinnate +, lyrate-pinnate +, 3-foliolate +  and simple +
opposite +  and alternate +
winter-persistent +
cauline +  and basal +
cordate +, reniform +, orbiculate +, oblanceolate +, obovate +, oblong +, elliptic +  and rhombic +
oblanceolate +  and obovate +
palmate +  and pinnate +
crenate +, dentate +, laciniate +, 2-7-lobed +, flat +, lobed +, toothed +  and entire +
inferior +  and superior +
clustered +, biseriate +  and superposed +
stipitate-glandular +  and puberulent +
purple-veined +
0 (?) +  and 4 (?) +
free +  and distinct +
suborbiculate;elliptic;suborbiculate;elliptic;obovate +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Flowering late spring–summer. +
adnate +, free +, connate +  and distinct +
lyrate-pinnate +  and pinnate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
not arillate +
persistent +
free +  and distinct +
reflexed +  and erect-spreading +
deltate-lanceolate +, deltate-ovate +  and deltate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
free +  and distinct +
hirsute;hirsute;puberulent +
1 +  and 5 +
30 cm300 mm <br />0.3 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
palmate +  and pinnate +
0.7 cm7 mm <br />0.007 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
ovate;lanceolate;ovate;lanceolate;linear +
0.3 cm3 mm <br />0.003 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
geniculate-jointed +  and entire +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
hairy +  and glabrous +
not hooked +  and hooked +
accrescent +
10 cm100 mm <br />0.1 m <br /> (?) +
G. macrophyllum subsp. perincisum +, G. macrophyllum var. rydbergii +, G. oregonense +  and G. perincisum var. intermedium +
Geum macrophyllum var. perincisum +
Geum macrophyllum +
variety +
usually much larger +
inconspicuous +
hemispheric +  and cylindric +
enlarged +
geniculate-jointed +
hairy +  and glabrous +