Eremogone congesta

(Nuttall) Ikonnikov

Novosti Syst. Vyssh. Rast. 10: 139. 1973.

Common names: Ballhead sandwort
Endemic
Basionym: Arenaria congesta Nuttall in J. Torrey and A. Gray, Fl. N. Amer. 1: 178. 1838
Treatment appears in FNA Volume 5. Treatment on page 60. Mentioned on page 57, 58, 61, 62.

Plants tufted or sometimes matted, green, not glaucous, with woody base. Stems ± erect, 3–40 (–50) cm, glabrous or often stipitate-glandular. Leaves: basal leaves persistent or not; cauline leaves in 3–5 pairs, similar, but reduced distally; basal blades erect-ascending to arcuate-spreading, subulate or needlelike to filiform, (0.8–) 2–11 (–14) cm × 0.4–2 mm, flexuous or rigid, herbaceous to ± fleshy, apex obtuse to sharply acute or spinose, glabrous, sometimes glaucous. Inflorescences 3–50+-flowered, congested and capitate or sometimes open, umbellate cymes. Pedicels 0.1–7 (–15) mm or ± absent, usually glabrous, rarely stipitate-glandular. Flowers: sepals 1–3-veined, sometimes obscurely so, ovate to lanceolate, 3–6.5 mm, not expanding in fruit, margins narrow, apex obtuse or acute to acuminate, rarely spinose, glabrous (or glandular in var. prolifera); petals white, oblong, 5–8 (–10) mm, 1.5–2 times as long as sepals, apex entire to slightly emarginate; nectaries as lateral and abaxial mound with crescent-shaped groove at base of filaments opposite sepals, 0.3 × 0.15–0.2 mm. Capsules 3.5–6 mm, glabrous. Seeds reddish-brown to black, broadly ellipsoid to ovoid, 1.4–3 mm, tuberculate; tubercles low, rounded, often elongate.

Distribution

V5 120-distribution-map.gif

Alta., Sask., Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Varieties 9 (9 in the flora).

Eremogone congesta is highly polymorphic; it has been been divided into 11 varieties (nine recognized here), most of which are distinctive and locally distributed. M. F. Baad (1969) noted two patterns of variation of different origin within E. congesta, but he did not present a revised classification.

While most specimens of the four varieties with dense inflorescences do not exhibit evident pedicels, the occasional plant does bear one or more pedicels to 1–2 mm, sometimes in secondary inflorescences.

Selected References

None.

Key

1 Pedicels usually 0.1-0.2 mm or ± absent; inflorescences dense, tight cymes > 2
1 Pedicels 1-6(-15) mm; inflorescences somewhat to markedly open cymes or umbellate cymes > 5
2 Basal leaf blades filiform, 3-14 cm > 3
2 Basal leaf blades needlelike, 1-3.5 cm > 4
3 Inflorescences capitate, rounded cymes; sepal apices obtuse to rounded Eremogone congesta var. congesta
3 Inflorescences capitate, pyramidal cymes; sepal apices narrowly acute to acuminate Eremogone congesta var. cephaloidea
4 Sepals 5-6.5 mm, apex acute to acuminate; basal leaf blades herbaceous; California, Nevada Eremogone congesta var. simulans
4 Sepals 3.5-4 mm, apex obtuse; basal leaf blades ± succulent; California, Oregon Eremogone congesta var. crassula
5 Sepal apices obtuse > 6
5 Sepal apices acute to acuminate or spinose > 7
6 Inflorescences proliferating, ± loose cymes; bracts often not closely enveloping sepals; Alberta, Saskatchewan, Colorado, Idaho, Montana, Utah, Wyoming Eremogone congesta var. lithophila
6 Inflorescences umbels, bracts clustered at umbel base; California Eremogone congesta var. suffrutescens
7 Basal leaf blades (2-)3-8 cm, filiform; pedicels glabrous or sometimes stipitate-glandular (possible throughout range, true for Nevada populations); Idaho, Nevada, Oregon, Washington Eremogone congesta var. prolifera
7 Basal leaf blades 0.5-2.5(-3.5) cm, needlelike or filiform; pedicels glabrous; California, Nevada, Utah > 8
8 Inflorescences proliferating, ± loose cymes; sepals 3.5-4.5 mm Eremogone congesta var. subcongesta
8 Inflorescences capitate and often rounded, to subcongested, proliferating or open cymes; sepals 4.5-6.5 mm > 9
9 Sepals 4.5-5.5 mm, weakly to conspicuously 1-3-veined, apex spinose Eremogone congesta var. charlestonensis
9 Sepals usually 5.5-6.5 mm, conspicuously 3-veined, apex acute to acuminate Eremogone congesta var. simulans
... more about "Eremogone congesta"
entire +  and slightly emarginate +
glabrous +  and glaucous +
acute;acuminate +
Ronald L. Hartman +, Richard K. Rabeler +  and Frederick H. Utech +
(Nuttall) Ikonnikov +
11 cm110 mm <br />0.11 m <br /> (14 cm140 mm <br />0.14 m <br />) +
2 cm20 mm <br />0.02 m <br /> (11 cm110 mm <br />0.11 m <br />) +
needlelike +  and filiform +
herbaceous +  and more or less fleshy +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and persistent +
Arenaria congesta +
filiform +  and subulate or narrowly linear +
not +  and succulent +
reduced +
ovoid;urceolate +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Ballhead sandwort +
dish--shaped +
expanded +
Alta. +, Sask. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
straight +  and curved +
central +  and peripheral +
inconspicuous +
unisexual +  and bisexual +
stalked +  and simple +
crescent--shaped +
disc--shaped +  and dish +
open +  and 3-50+-flowered +
connate +
0.15mm;0.2mm +
swollen +
3 +  and 5 +
crassinucellate +  and bitegmic +
campylotropous +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
stipitate-glandular +  and glabrous +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
fugacious +
1.5-2 times as long as sepals +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Novosti Syst. Vyssh. Rast. +
reddish-brown;black +
tuberculate +
broadly ellipsoid +  and ovoid +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
persistent +
connate +  and distinct +
ovate;lanceolate +
not expanding +
0.3 cm3 mm <br />0.003 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
branched +  and simple +
40 cm400 mm <br />0.4 m <br /> (50 cm500 mm <br />0.5 m <br />) +
stipitate-glandular +  and glabrous +
3 cm30 mm <br />0.03 m <br /> (40 cm400 mm <br />0.4 m <br />) +
[50 +  and 3 +
smooth +  and papillate +
distinct +
filiform +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
stout +  and slender +
Eremogone congesta +
Eremogone +
species +
ascending;recurved +
divided +
elongate +  and rounded +
10 +  and 8 +
rhizomatous +  and taprooted +
plant +  and matted +