Drymocallis pseudorupestris

(Rydberg) Rydberg

Monogr. N. Amer. Potentilleae, 194. 1898.

Endemic
Basionym: Potentilla pseudorupestris Rydberg Bull. Torrey Bot. Club 24: 250, plate 307. 1897
Synonyms: Drymocallis glandulosa subsp. pseudorupestris (Rydberg) Soják P. glandulosa subsp. pseudorupestris (Rydberg) D. D. Keck P. glandulosa var. pseudorupestris (Rydberg) Breitung P. rupestris var. americana Th. Wolf
Treatment appears in FNA Volume 9. Treatment on page 289. Mentioned on page 283, 284, 287, 288, 290.

Caudex branches elongate. Stems openly tufted to loosely spaced, (0.3–) 0.6–4 dm; base 1–3 mm diam., ± densely septate-glandular. Leaves glabrate or sparsely to ± densely hairy; basal (2–) 3–16 cm, leaflet pairs (2–) 3–4 (–5); terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3 (–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse; cauline 0–2, reduced, leaflet pairs 2–3. Inflorescences 2–40-flowered, not or ± leafy, open, 1/6–3/4 (–4/5) of stem, ± wide, branch angles (10–) 20–40 (–50) °. Pedicels 3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular. Flowers opening widely; epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm; sepals spreading, 4–7 (–9) mm, apex acute to obtuse, apiculate; petals overlapping or not, spreading, cream-white to pale-yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals; filaments 1–4 mm, anthers 0.7–1.2 mm; styles thickened, 1–1.5 mm. Achenes light-brown, 1 mm.

Distribution

V9 465-distribution-map.jpg

Alta., B.C., Calif., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Varieties 3 (3 in the flora).

Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range.

Selected References

None.

Key

1 Stems (1–)2–4 dm, bases (1.5–)2–3 mm diam.; basal leaves (4–)7–16 cm; terminal leaflets (1–)2–3(–4) cm, teeth usually double, (5–)8–15 per side; flowers (5–)10–40; petals 6–12 × 5–11 mm, widely overlapping; filaments 2–4 mm. Drymocallis pseudorupestris var. pseudorupestris
1 Stems (0.3–)0.6–2.5 dm, bases 1–2(–3) mm diam.; basal leaves (2–)3–9(–15) cm; terminal leaflets 0.2–2(–4) cm, teeth single or ± double, 2–8(–12) per side; flowers 2–12(–20); petals 4–8(–9) × 3–6(–8) mm, not or ± overlapping; filaments 1–2.5(–3) mm > 2
2 Basal leaves: leaflet pairs (2–)3(–4); hypanthia and sepals not bristly or bristles less than 1 mm; short hairs sparse to moderately abundant on stems and pedicels (sometimes absent); styles usually golden brown, rarely reddish. Drymocallis pseudorupestris var. saxicola
2 Basal leaves: leaflet pairs 3–4(–5); hypanthia and sepals prominently bristly, bristles 1–1.5 mm; short hairs absent or sparse on stems and pedicels; styles usually dark red, rarely golden brown. Drymocallis pseudorupestris var. crumiana

"thin" is not a number."wide" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

light-brown +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
acute;obtuse +
Barbara Ertter +
(Rydberg) Rydberg +
compound +  and simple +
opposite +  and alternate +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
3 cm30 mm <br />0.03 m <br /> (16 cm160 mm <br />0.16 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Potentilla pseudorupestris +
oblanceolate-oblong +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
(10-)20-40(-50)° +
free +  and distinct +
Alta. +, B.C. +, Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
not +  and aggregated +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
less;10;80 +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
open +, leafy +  and 2-40-flowered +
1/6 +  and 3/4-4/5 +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
hairy +  and glabrate +
overlapping +  and not confluent +
elliptic-ovate +  and flabellate or rhombic +
toothed +, or +, flat +, entire +  and more or less toothed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (2 cm20 mm <br />0.02 m <br />) +
not +  and overlapping +
0 (?) +  and 4 (?) +
cream-white +  and pale-yellow +
free +  and distinct +
4mm;12mm +
obovate +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
adnate +, free +, connate +  and distinct +
Monogr. N. Amer. Potentilleae, +
not arillate +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
persistent +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
free +  and distinct +
branched +  and simple +
openly tufted +  and loosely spaced +
stramineous +, reddish +  and green +
1 +  and 20 +
palmate +  and pinnate +
narrowly;broadly-triangular-ovate +
deciduous +
distinct +
fusiform +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Drymocallis glandulosa subsp. pseudorupestris +, P. glandulosa subsp. pseudorupestris +, P. glandulosa var. pseudorupestris +  and P. rupestris var. americana +
Drymocallis pseudorupestris +
Drymocallis +
species +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (3 cm30 mm <br />0.03 m <br />) +
broadly obovate-cuneate +  and flabellate +
0.5 cm5 mm <br />0.005 m <br /> (3 cm30 mm <br />0.03 m <br />) +
horseshoe--shaped +
2 (?) +  and 15 (?) +
inconspicuous +
hemispheric +  and conic +
enlarged +
peglike-glandular +  and septate-glandular +