Drymocallis fissa

(Nuttall) Rydberg

Monogr. N. Amer. Potentilleae, 197. 1898.

Common names: Leafy drymocallis or wood beauty
EndemicIllustrated
Basionym: Potentilla fissa Nuttall in J. Torrey and A. Gray, Fl. N. Amer. 1: 446. 1840
Treatment appears in FNA Volume 9. Treatment on page 284. Mentioned on page 282, 286, 290.

Caudex branches short-to-elongate. Stems tufted to loosely spaced, (1.2–) 1.5–3.5 (–4.5) dm; base (1.5–) 2–3 mm diam., ± densely septate-glandular. Leaves sparsely to moderately hairy; basal (3–) 7–19 cm, leaflet pairs (4–) 5–6 (–10; additional reduced leaflets sometimes interspersed); terminal leaflet usually broadly obovate-cuneate, sometimes elliptic, (1–) 1.5–3.5 (–5) × (1–) 1.5–3 (–3.5) cm, teeth single or double, 5–13 per side, apex rounded to obtuse; cauline 1–3, well developed, leaflet pairs 4–6 (–10). Inflorescences 5–15-flowered, leafy, congested to ± open, 1/6–1/2 of stem, narrow to wide, branch angles 15–30 (–40) °. Pedicels 1–12 mm, short-hairy, septate-glandular. Flowers opening widely; epicalyx bractlets linear-oblanceolate, 3–7 × 1–2 mm; sepals spreading, 6–10 mm, apex acute to acuminate; petals overlapping, spreading, yellow, broadly obovate, 7–11 × 5–11 mm, equal to or exceeding sepals; filaments 1.5–4.5 mm, anthers (0.7–) 1–1.4 mm; styles thickened, 1 mm. Achenes light-brown, 1 mm.


Phenology: Flowering May–Aug.
Habitat: Sagebrush slopes, open forests, stream banks, often in rocky or moderately disturbed sites
Elevation: 1600–3000 m

Discussion

Drymocallis fissa is distinctive in its relatively numerous leaflets (often with additional smaller ones), large flowers, and large, elongate anthers. It is most abundant in the Colorado Front Range, extending into the Medicine Bow Mountains of Wyoming. Outlying populations occur at least as far north as the Bighorn Mountains of northern Wyoming and the Black Hills of South Dakota. Tentatively included here are large-anthered populations from the eastern Uintah Mountains of Utah, though these often have fewer leaflets and smaller flowers of unknown color; they may represent a unique taxon worthy of separate recognition. Possible collections of D. fissa from New Mexico, including the type of Potentilla fissa var. major Torrey & A. Gray, are of uncertain placement in that they combine features of D. arguta, D. arizonica, and D. fissa. Reports from other states, including Montana, are probably all based on misidentified specimens.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Drymocallis fissa"
light-brown +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
acute;acuminate +
Barbara Ertter +
(Nuttall) Rydberg +
compound +  and simple +
opposite +  and alternate +
3 cm30 mm <br />0.03 m <br /> (7 cm70 mm <br />0.07 m <br />) +
7 cm70 mm <br />0.07 m <br /> (19 cm190 mm <br />0.19 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Potentilla fissa +
oblanceolate-oblong +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
short +  and elongate +
Leafy drymocallis or wood beauty +
Colo. +, S.Dak. +, Utah +  and Wyo. +
not +  and aggregated +
1600–3000 m +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
linear-oblanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
less;10;80 +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Sagebrush slopes, open forests, stream banks, often in rocky or moderately disturbed sites +
congested +  and more or less open +
1/6 +  and 1/2 +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
overlapping +  and not confluent +
elliptic-ovate +  and flabellate or rhombic +
toothed +, or +, flat +, entire +  and more or less toothed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.1 cm1 mm <br />0.001 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
overlapping +
0 (?) +  and 4 (?) +
free +  and distinct +
7mm;11mm +
obovate +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Flowering May–Aug. +
adnate +, free +, connate +  and distinct +
Monogr. N. Amer. Potentilleae, +
not arillate +
persistent +
free +  and distinct +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Endemic +  and Illustrated +
free +  and distinct +
branched +  and simple +
tufted +  and loosely spaced +
stramineous +, reddish +  and green +
1 +  and 20 +
narrow +  and wide +
palmate +  and pinnate +
narrowly;broadly-triangular-ovate +
deciduous +
distinct +
fusiform +
1 cm10 mm <br />0.01 m <br /> (?) +
Closterostyles +, Potentilla sect. Closterostyles +, Potentilla subg. Closterostyles +  and Potentilla sect. Rupestres +
Drymocallis fissa +
Drymocallis +
species +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
obovate-cuneate +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
horseshoe--shaped +
5 (?) +  and 13 (?) +
inconspicuous +
hemispheric +  and conic +
enlarged +
peglike-glandular +  and septate-glandular +