Draba lactea

Adams

Mém. Soc. Imp. Naturalistes Moscou 5: 104. 1817.

Synonyms: Draba allenii Fernald Draba boecheri Gjaerevoll & Ryvarden Draba fernaldiana Polunin
Treatment appears in FNA Volume 7. Treatment on page 316. Mentioned on page 270, 273, 275, 279, 280, 286.

Perennials; (cespitose); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose. Stems unbranched, 0.2–1.1 (–1.5) dm,glabrous throughout or sparsely pubescent proximally, trichomes short-stalked, substellate, 2–8-rayed, (non-crisped), 0.5–0.3 mm. Basal leaves rosulate; petiolate; petiole (persistent, strongly thickened), margin usually ciliate, (trichomes usually simple and 2-rayed, 0.3–1 mm); blade oblanceolate to obovate, (0.3–) 0.5–1.1 (–1.7) cm × (1–) 2–6 mm, margins usually entire, rarely denticulate, (sometimes ciliate), surfaces sometimes pubescent with stellate to subdendritic, 4–12-rayed, (non-crisped) trichomes, 0.1–0.4 mm, (midvein persistent, prominent, strongly thickened). Cauline leaves 0 (or, rarely, 1 as a bract). Racemes 2–8 (–12) -flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent as stem basally. Fruiting pedicels divaricate-ascending, straight, (1–) 2–5 (–10) mm, glabrous. Flowers: sepals ovate, 1.8–3 mm, usually glabrous, rarely sparsely pubescent subapically, (trichomes simple); petals white, obovate, 3–5 × 1.8–3 mm; anthers ovate, 0.3–0.4 mm. Fruits oblong to elliptic-lanceolate or ovate to broadly so, plane, flattened, 4–8 × (1.5–) 2–3 mm; valves glabrous; ovules (10–) 14–22 (–26) per ovary; style 0.1–0.4 mm. Seeds ovoid, 0.8–1.1 × 0.5–0.6 mm. 2n = 32, 48.


Phenology: Flowering Jun–Aug.
Habitat: Rock outcrops, talus, rocky hillsides and ridges, open gravelly areas, seepage swales, meadows
Elevation: 0-2000 m

Distribution

V7 421-distribution-map.gif

Greenland, Nfld. and Labr., N.W.T., Nunavut, Que., Yukon, Alaska, Europe (n Finland), Europe (Norway), Europe (w Sweden), Asia (Russian Far East), Asia (c), Asia (n Siberia), Atlantic Islands (Iceland), circumpolar

Discussion

T. W. Böcher (1966) postulated that Draba lactea originated from hybridization between D. fladnizensis and D. nivalis, but A.-C. Scheen et al. (2002) showed that it is more closely allied to D. subcapitata. By contrast, H. H. Grundt et al. (2004) concluded that hexaploid D. lactea originated from tetraploids of the same species, which in turn originated from the diploid D. palanderiana lineage. They suggested that D. lactea probably originated multiple times in the Beringian area and migrated to reach its present circumpolar distribution. The hexaploids are distributed throughout the species range, whereas the tetraploids are known only from Alaska and the Russian Far East (Grundt et al. 2005b).

Draba fernaldiana, which was collected from Southampton Island (Nunavut), was not mentioned by R. C. Rollins (1993). The plants are completely glabrous except for leaf margins, which are ciliate with simple and sparse 2-rayed trichomes. The taxon resembles some forms of D. lactea and is tentatively herein included within that species. The only conflict in such placement is petal color, which was listed in the original description of D. fernaldiana as pale yellow instead of white.

Glabrous or glabrescent forms of Draba lactea are quite common in the Canadian Arctic Archipelago, whereas pubescent forms predominate in Alaska and the Russian Far East.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Draba lactea"
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
Ihsan A. Al-Shehbaz +, Michael D. Windham +  and Reidar Elven +
appendaged +  and unappendaged +
1.1 cm11 mm <br />0.011 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and auriculate +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
lobed +, toothed +  and entire +
oblanceolate;obovate +
much smaller +  and reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
urceolate +, campanulate +  and tubular +
scapose +  and branched +
sessile +  and petiolate +
well-developed +
distinct +
well-differentiated +
emarginate +  and entire +
Greenland +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Que. +, Yukon +, Alaska +, Europe (n Finland) +, Europe (Norway) +, Europe (w Sweden) +, Asia (Russian Far East) +, Asia (c) +, Asia (n Siberia) +, Atlantic Islands (Iceland) +  and circumpolar +
0-2000 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
not basally +  and dilated +
actinomorphic +
orange;purple;orange;purple;pink;white;yellow +
spreading +, ascending +  and erect +
linear +, oblanceolate +, spatulate +, obovate +, subsaccate +, not saccate +, oblong +  and ovate +
latiseptate +, twisted +  and sessile +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
indehiscent +  and dehiscent +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
flattened +, plane +, oblong +  and elliptic-lanceolate or ovate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Rock outcrops, talus, rocky hillsides and ridges, open gravelly areas, seepage swales, meadows +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
denticulate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
rudimentary +
obovate +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Jun–Aug. +
trinucleate +  and 3(-11)-colpate +
Mém. Soc. Imp. Naturalistes Moscou +
ebracteate;2-8(-12)-flowered +
ascending +  and erect +
slender +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
straight +  and not flexuous +
divaricate-ascending +
glabrous +  and pubescent +
mucilaginous +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
pubescent +  and glabrous +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
reduced +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
tetradynamous +
procumbent +  and decumbent +
prostrate +  and decumbent +
usually erect +  and ascending +
pubescent +  and glabrous +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
Draba allenii +, Draba boecheri +  and Draba fernaldiana +
Draba lactea +
species +
subdendritic +  and short-stalked +
substellate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
scapose +  and not +
pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +