Cyperus squarrosus

Linnaeus

Cent. Pl. II, 6. 1756.

Illustrated
Synonyms: Chlorocyperus inflexus (Muhlenberg) Palla Cyperus aristatus Rottbøll Cyperus aristatus var. inflexus (Muhlenberg) Kükenthal Cyperus aristatus var. runyonii O’Neill Cyperus inflexus (Linnaeus) C. B. Clarke Dichostylis aristata Mariscus squarrosus
Treatment appears in FNA Volume 23. Treatment on page 165. Mentioned on page 144, 145, 166.

Herbs, annual, cespitose, with fibrous-roots. Culms 1–20, trigonous, 2–16 cm × 0.5–0.8 mm, glabrous. Leaves 1–3, flat to V-shaped, (1–) 5–10 (–15) × 0.5–2.5 mm. Inflorescences: spike 1, loosely to densely ovoid to oblong, 6–20 (–40) × 9–15 (–20) mm; rays absent or 1–3 (–6), 4–40 mm; bracts (1–) 2–4, longest ± erect, V-shaped, 1–15 cm × 0.5–3 mm; rachilla ± deciduous, wingless. Spikelets (2–) 6–20 (–40), greenish to reddish-brown, somewhat compressed, ovoidlanceoloid to oblong, 2.5–10 (–20) × 1.3–2.2 mm (excluding awns); floral scales deciduous, (4–) 10–20 (–34), greenish to stramineous or brownish red laterally, greenish medially, (5–) 7–9 (–11) -ribbed nearly to margins, oblong-lanceolate, (1.2–) 1.3–1.8 (–2.2) × (0.5–) 0.7–0.8 (–1) mm, apex cuspidate, excurved awn additional 0.5–1 (–1.3) mm. Flowers: stamen 1, filaments 1.5 mm; anthers oblong, 0.2–0.3 mm, connective apex reddish, minute; styles 0.3–0.5 mm; stigmas 0.4–0.7 mm. Achenes light-brown to nearly black, stipitate, obovoid (occasionally linear-spatulate or linear-oblong, infrequently constricted in middle), 0.7–0.8 (–1.1) × (0.2–) 0.3–0.4 (–0.5) mm, stipe 0.05–0.1 × 0.1 mm, apex truncate, apiculate, surfaces papillose.


Phenology: Fruiting summer.
Habitat: Moist, disturbed soils, gravelly roadsides, flood plains, edges of puddles, muddy places
Elevation: 0–2500 m

Distribution

V23 265-distribution-map.jpg

Alta., B.C., Man., N.B., Ont., Que., Ala., Ariz., Ark., Calif., Colo., Conn., Del., D.C., Fla., Ga., Idaho, Ill., Ind., Iowa, Kans., Ky., La., Maine, Md., Mass., Mich., Minn., Miss., Mo., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.C., S.Dak., Tenn., Tex., Utah., Vt., Va., Wash., W.Va., Wis., Wyo., Mexico, West Indies, Bermuda, Central America, South America, Eurasia, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia

Discussion

Cyperus squarrosus can be recognized by its small size and annual habit combined with its oblong-lanceolate floral scales bearing five to eleven conspicuous ribs and excurved awns. Some collections have been misidentified as C. acuminatus, an annual species of subg. Pycnostachys that has ovate-lanceolate, three-ribbed floral scales and digitately clustered spikelets.

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Cyperus squarrosus"
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
light-brown +  and nearly black +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
0.3mm;0.4mm +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
apiculate +, truncate +  and cuspidate +
Gordon C. Tucker* +, Brian G. Marcks* +  and J. Richard Carter * +
Linnaeus +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
additional +
excurved +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +  and deciduous +
decurrent +
w--shaped +  and keeled +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
1 cm10 mm <br />0.01 m <br /> (15 cm150 mm <br />0.15 m <br />) +
v--shaped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
parallel +  and divergent +
terete +, rolled +  and plicate +
2 cm20 mm <br />0.02 m <br /> (16 cm160 mm <br />0.16 m <br />) +
scabridulous +
trigonous +
0.5mm;0.8mm +
Alta. +, B.C. +, Man. +, N.B. +, Ont. +, Que. +, Ala. +, Ariz. +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Fla. +, Ga. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, La. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Miss. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.C. +, S.Dak. +, Tenn. +, Tex. +, Utah. +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +, Wyo. +, Mexico +, West Indies +, Bermuda +, Central America +, South America +, Eurasia +, Africa +, Atlantic Islands +, Indian Ocean Islands +, Pacific Islands +  and Australia +
0–2500 m +
1.5 cm15 mm <br />0.015 m <br /> (?) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
greenish +  and stramineous or brownish red laterally +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
oblong-lanceolate +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
hypogynous +  and subtending +
biconvex +  and trigonous +
Moist, disturbed soils, gravelly roadsides, flood plains, edges of puddles, muddy places +
subumbellate +  and capitate +
spreading +  and erect +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (?) +  and 1 cm10 mm <br />0.01 m <br /> (?) +
flat +  and v-shaped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting summer. +
2-3(-4)-carpellate +
antrorse +  and extrorse +
3 (?) +  and 1 (?) +
Cent. Pl. II, +
conspicuous +
unequal +
1 +  and 3 +
0.4 cm4 mm <br />0.004 m <br /> (4 cm40 mm <br />0.04 m <br />) +
adventitious +
basal +  and proximal +
cylindric;compressed +
cylindric +
W2 +  and Illustrated +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
ovoid;oblong +
0.9 cm9 mm <br />0.009 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
greenish +  and reddish-brown +
0.25 cm2.5 mm <br />0.0025 m <br /> (1 cm10 mm <br />0.01 m <br />) +
6 +  and 20 +
oblong;compressed +
1.3mm;2.2mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (?) +  and 0.01 cm0.1 mm <br />1.0e-4 m <br /> (?) +
0.1 cm1 mm <br />0.001 m <br /> (?) +
2-3-fid +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
papillose +
Chlorocyperus inflexus +, Cyperus aristatus +, Cyperus aristatus var. inflexus +, Cyperus aristatus var. runyonii +, Cyperus inflexus +, Dichostylis aristata +  and Mariscus squarrosus +
Cyperus squarrosus +
Cyperus subg. Cyperus +
species +
tuberous +, stoloniferous +  and rhizomatous +
herb +  and cespitose +