Cyperus sanguinolentus

Vahl

Enum. Pl. 2: 351. 1805.

IntroducedIllustrated
Synonyms: Cyperus louisianensis Thieret Pycreus sanguinolentus (Vahl) Nees
Treatment appears in FNA Volume 23. Treatment on page 161. Mentioned on page 142, 162.

Herbs, perennial; rhizomes ± horizontal, to 12 cm × 0.8–1.2 mm. Culms ± terete to roundly trigonous, 3–25 (–60) cm × 0.3–2 mm, glabrous. Leaves 1–3, V-shaped, 1–8 (–15) cm × 1–2 mm. Inflorescences: spike 1, loosely ovoid, 7–14 × 8–20 mm; rays (0–) 1–4, to 2 (–3) cm; bracts 2–4, horizontal to ascending at 30 (–45) °, V-shaped to flat, 1–8 (–18) cm × 0.5–1.5 (–2) mm. Spikelets 3–5 (–14), oblong to linear-lanceoloid, 8–18 × 2–2.6 (–3) mm; floral scales (6–) 10–26 (–32), laterally clear to light-brown, sometimes with purplish margins, medially light-brown, 2-keeled, laterally ribless, medially 2–3-ribbed, oblong to ovate, 1.9–2.2 (–2.7) × 1.8–2.3 mm, apex obtuse. Flowers: stamens 3; anthers ellipsoid, 0.3–0.6 mm, connective not prolonged; styles 0.6–1 mm; stigmas 1–1.5 mm. Achenes brown, ± stipitate, obovoid to ovoid, 1–1.4 × 0.6–0.8 mm, apex truncate, apiculate, surfaces minutely punctate.


Phenology: Fruiting late summer–fall.
Habitat: Emergent shorelines, ditches
Elevation: 20 m

Distribution

V23 257-distribution-map.jpg

Introduced; Ala., Ga., La., Miss., Asia, Africa

Discussion

Cyperus sanguinolentus is the only rhizomatous, distigmatic Cyperus in the eastern United States. An early collection from Louisiana was described as C. louisianensis, supposedly endemic. Recent field work and morphometric studies convincingly showed it to be an introduction of the Asian C. sanguinolentus) (J. R. Carter and C. T. Bryson 2001.

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Cyperus sanguinolentus"
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
obovoid;ovoid +
0.6mm;0.8mm +
ellipsoid +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
apiculate +  and truncate +
Gordon C. Tucker* +, Brian G. Marcks* +  and J. Richard Carter * +
persistent +  and deciduous +
decurrent +
Undefined subg. Pycreus +
w--shaped +  and keeled +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
8 cm80 mm <br />0.08 m <br /> (18 cm180 mm <br />0.18 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1 cm10 mm <br />0.01 m <br /> (8 cm80 mm <br />0.08 m <br />) +
horizontal +  and ascending +
v-shaped +  and flat +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
parallel +  and divergent +
terete +, rolled +  and plicate +
not prolonged +
25 cm250 mm <br />0.25 m <br /> (60 cm600 mm <br />0.6 m <br />) +
3 cm30 mm <br />0.03 m <br /> (25 cm250 mm <br />0.25 m <br />) +
scabridulous +
less terete +  and roundly trigonous +
0.3mm;2mm +
Ala. +, Ga. +, La. +, Miss. +, Asia +  and Africa +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
light-brown +  and laterally clear +
0.19 cm1.9 mm <br />0.0019 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
oblong +  and ovate +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
hypogynous +  and subtending +
biconvex +  and trigonous +
Emergent shorelines, ditches +
subumbellate +  and capitate +
spreading +  and erect +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
8 cm80 mm <br />0.08 m <br /> (15 cm150 mm <br />0.15 m <br />) +
1 cm10 mm <br />0.01 m <br /> (?) +  and 8 cm80 mm <br />0.08 m <br /> (?) +
v--shaped +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting late summer–fall. +
2-3(-4)-carpellate +
antrorse +  and extrorse +
3 (?) +  and 1 (?) +
nearly +  and wingless +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
unequal +
1 +  and 4 +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0 cm0 mm <br />0 m <br /> (?) +  and 12 cm120 mm <br />0.12 m <br /> (?) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (?) +  and 0.12 cm1.2 mm <br />0.0012 m <br /> (?) +
adventitious +
basal +  and proximal +
cylindric;compressed +
cylindric +
Introduced +  and Illustrated +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
loosely spicate +  and more or less digitate +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
oblong;linear-lanceoloid +
2mm;2.6mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
2-3-fid +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Cyperus louisianensis +  and Pycreus sanguinolentus +
Cyperus sanguinolentus +
Cyperus subg. Pycreus +
species +
tuberous +, stoloniferous +  and rhizomatous +