Cardamine pensylvanica

Muhlenberg ex Willdenow

Sp. Pl. 3: 486. 1801.

WeedyEndemic
Synonyms: Cardamine breweri var. oregana (Piper) Detling Cardamine flexuosa subsp. pensylvanica (Muhlenberg ex Willdenow) O. E. Schulz Cardamine hirsuta var. pensylvanica (Muhlenberg ex Willdenow) P. W. Graff Cardamine multifolia Rydberg Cardamine oregana Piper Cardamine pensylvanica var. brittoniana Farwell Cardamine rotundifolia var. diversifolia O. E. Schulz Dracamine pensylvanica
Treatment appears in FNA Volume 7. Treatment on page 482. Mentioned on page 466.

Annuals or biennials; sparsely hirsute basally, glabrous distally. Rhizomes absent. Stems (simple from base), erect, (not flexuous), unbranched or branched distally, (0.5–) 1.5–5.5 (–7) dm. Basal leaves (soon withered), not rosulate, similar to proximalmost cauline leaves, 4–15 cm. Cauline leaves (3–) 5–20 (–35), pinnately (5 or) 7–13 (–19) -foliolate, sometimes appearing pinnatisect, lobe number similar to leaflets (middle and proximal ones 2–11 cm), petiolate, leaflets petiolulate, subsessile, or sessile; petiole (0.4–) 1–3.5 (–4.5) cm, base not auriculate, (often sparsely hirsute); lateral leaflets shortly petiolulate or sessile, (decurrent on rachis smaller than terminal, distalmost blades narrower, with fewer lobes or leaflets), margins entire or crenate; terminal leaflet (subsessile or petiolule to 1 cm), blade suborbicular, obovate to oblanceolate, or elliptic, 1.3–3 (–4) cm × 6–25 mm, base often cuneate, margins entire, repand, or obscurely 3 or 5-lobed. Racemes ebracteate. Fruiting pedicels divaricate-ascending, (3–) 4–10 (–13) mm. Flowers: sepals oblong, (1–) 1.3–2.3 × 0.5–1 mm, lateral pair not saccate basally; petals white, narrowly spatulate to oblanceolate, 2–3.5 (–4) × 0.8–1.5 mm, (not clawed); filaments: median pairs 1.5–2.5 mm, lateral pair 1–2 mm; anthers ovate, 0.2–0.3 mm. Fruits linear, (torulose), (1.4–) 1.7–2.7 (–3.2) cm × 0.8–1.1 mm; ovules 40–80 per ovary; style 0.5–1 mm. Seeds brown, oblong to ovoid, 0.7–1.1 × 0.5–0.8 mm. 2n = 32, 64.


Phenology: Flowering Apr–Jul.
Habitat: Marshes, streams, swamps, ditches, seepage, springs, lake margins, mesic bottomland and upland forests, wet areas, ledges of sheltered bluffs, banks and shallow water of streams and spring branches, margins of crop fields, waste ground
Elevation: 0-2800 m

Distribution

V7 743-distribution-map.gif

Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Ont., P.E.I., Que., Sask., Yukon, Ala., Alaska, Ark., Calif., Colo., Conn., Del., D.C., Fla., Ga., Idaho, Ill., Ind., Iowa, Kans., Ky., La., Maine, Md., Mass., Mich., Minn., Miss., Mo., Mont., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.C., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wis., Wyo.

Discussion

We have not seen material of Cardamine pensylvanica from Nebraska or South Dakota; it is very likely that the species grows in these states as well.

Selected References

None.

Lower Taxa

None.

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... more about "Cardamine pensylvanica"
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
Ihsan A. Al-Shehbaz +, Karol Marhold +  and Judita Lihová +
Muhlenberg ex Willdenow +
4 cm40 mm <br />0.04 m <br /> (15 cm150 mm <br />0.15 m <br />) +
cuneate +  and not auriculate +
appendaged +  and unappendaged +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (3 cm30 mm <br />0.03 m <br />) +
compound +  and trifoliolate +
lobed +, 1-3-pinnatisect +, toothed +  and entire +
obovate;oblanceolate or elliptic +
much smaller +  and reduced +
0.6 cm6 mm <br />0.006 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and 7-13(-19)-foliolate +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Ala. +, Alaska +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Fla. +, Ga. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, La. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Miss. +, Mo. +, Mont. +, Nev. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.C. +, Tenn. +, Tex. +, Utah +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +  and Wyo. +
0-2800 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
lateral +  and median +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
actinomorphic +
not basally +  and saccate +
lilac;purple;lilac;purple;pink;white +
ascending +, spreading +  and erect +
oblanceolate +, spatulate +, obovate +, oblong +  and ovate +
latiseptate +  and unsegmented +
2.7 cm27 mm <br />0.027 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
angustiseptate +, 4-angled +  and terete +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
latiseptate +  and sessile +
torulose +, lanceolate +, oblong +  and linear +
Marshes, streams, swamps, ditches, seepage, springs, lake margins, mesic bottomland and upland forests, wet areas, ledges of sheltered bluffs, banks and shallow water of streams and spring branches, margins of crop fields, waste ground +
semiannular +  and annular +
sessile +  and petiolulate +
compound +, pinnately +, trifoliolate +  and simple +
alternate +, not +  and rosulate +
basal +  and cauline +
sessile +, subsessile +  and petiolulate +
connate +  and distinct +
decurrent +
5-lobed;repand;5-lobed;repand;entire;crenate;entire +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
2mm;3.5mm +
rudimentary +
narrowly spatulate +  and oblanceolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Flowering Apr–Jul. +
trinucleate +  and 3(-11)-colpate +
1 cm10 mm <br />0.01 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
divaricate-ascending +
stout +  and slender +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
flattened +
not +  and mucilaginous +
winged +  and margined +
uniseriate +  and biseriate +
not +  and mucilaginous +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
oblong +  and ovoid +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
persistent +  and caducous +
distinct +
erect +  and spreading or ascending +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
reduced +
latiseptate +  and unsegmented +
angustiseptate +, 4-angled +  and terete +
latiseptate +  and unsegmented +
angustiseptate +, 4-angled +  and terete +
Weedy +  and Endemic +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Cardamine breweri var. oregana +, Cardamine flexuosa subsp. pensylvanica +, Cardamine hirsuta var. pensylvanica +, Cardamine multifolia +, Cardamine oregana +, Cardamine pensylvanica var. brittoniana +, Cardamine rotundifolia var. diversifolia +  and Dracamine pensylvanica +
Cardamine pensylvanica +
Cardamine +
species +
scale-like +  and forked +
pubescent +, , +  and glabrous +
anastomosing +
biennial +  and annual +
hirsute +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +