Boechera pendulina

(Greene) W. A. Weber

Phytologia 51: 370. 1982.

Basionym: Arabis pendulina Greene Leafl. Bot. Observ. Crit. 2: 81. 1910
Synonyms: Arabis demissa var. russeola Rollins Arabis diehlii M. E. Jones Arabis pendulina var. russeola (Rollins) Rollins Arabis setulosa (Greene) W. A. Weber Boechera demissa var. pendulina (Greene) N. H. Holmgren Boechera demissa var. russeola (Rollins) N. H. Holmgren Boechera pendulina var. russeola (Rollins) Dorn
Treatment appears in FNA Volume 7. Treatment on page 395. Mentioned on page 352, 359, 377, 393, 400, 405.

Perennials; short to long-lived; sexual or apomictic; caudex sometimes woody. Stems usually 2–6 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.6–3 (–3.7) dm, sparsely to densely pubescent proximally, trichomes simple, 0.3–0.8 mm, glabrous distally. Basal leaves: blade oblanceolate to obovate, 1.5–6 mm wide, margins usually entire, rarely dentate, ciliate throughout, trichomes (usually simple), 0.4–1 mm, surfaces pubescent, trichomes simple and short and long-stalked, 2-rayed, 0.3–0.8 mm. Cauline leaves: 2–10 (–13), not concealing stem; blade auricles absent or, rarely, to 0.7 mm, surfaces of distalmost leaves glabrous or margins sparsely ciliate. Racemes 4–14-flowered, usually unbranched. Fruiting pedicels divaricate-ascending to horizontal, curved or angled downward, 3–7 (–10) mm, usually glabrous, rarely with some simple trichomes. Flowers divaricate-ascending at anthesis; sepals usually sparsely pubescent, rarely glabrous; petals whitish to pale lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid or spheroid. Fruits widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 2.2–4 cm × 1.2–2.1 mm; valves glabrous; ovules 40–70 (–90) per ovary; style 0.1–0.3 (–0.5) mm. Seeds biseriate, 0.9–1.2 × 0.6–0.9 mm, usually not winged. 2n = 14, 21.


Phenology: Flowering Apr–Jun.
Habitat: Rock outcrops, open gravelly flats and hillsides in sagebrush, pinyon-juniper, mountain mahogany, open conifer forests
Elevation: 1600-3100 m

Distribution

V7 564-distribution-map.gif

Ariz., Calif., Colo., Nev., Utah, Wyo.

Discussion

Although included in Arabis (Boechera) demissa by some authors (e.g., S. L. Welsh et al. 2003; N. H. Holmgren 2005b), B. pendulina is readily distinguished from that species (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). Typical collections are sexual diploids, whereas the type of var. russeola is a triploid apomict; further study is needed to determine if the two are conspecific.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera pendulina"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Greene) W. A. Weber +
not +  and rosulate +
Arabis pendulina +
appendaged +  and unappendaged +
0 cm0 mm <br />0 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate;obovate +
much smaller +  and reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Ariz. +, Calif. +, Colo. +, Nev. +, Utah +  and Wyo. +
2.2 cm22 mm <br />0.022 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
1600-3100 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
divaricate-ascending +
curved +  and nearly straight +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Rock outcrops, open gravelly flats and hillsides in sagebrush, pinyon-juniper, mountain mahogany, open conifer forests +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
whitish +  and pale lavender +
rudimentary +
oblanceolate +  and spatulate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Flowering Apr–Jun. +
trinucleate +  and 3(-11)-colpate +
spheroid +  and ellipsoid +
unbranched;4-14-flowered +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
orbicular +, oblong +  and flattened +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
glabrous +  and pubescent +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
2 +  and 6 +
subsessile +, sessile +  and petiolate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
persistent +
distinct +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
glabrous +  and pubescent +
Arabis demissa var. russeola +, Arabis diehlii +, Arabis pendulina var. russeola +, Arabis setulosa +, Boechera demissa var. pendulina +, Boechera demissa var. russeola +  and Boechera pendulina var. russeola +
Boechera pendulina +
Boechera +
species +
simple +  and long-stalked +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
downward +, divaricate-ascending +  and horizontal +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and long-lived +
hispid +, hirsute +, pubescent +  and glabrous +
apomictic +  and sexual +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +