Boechera inyoensis

(Rollins) Al-Shehbaz

Novon 13: 386. 2003.

Basionym: Arabis inyoensis Rollins Rhodora 43: 457. 1941
Synonyms: Arabis holboellii var. derensis S. L. Welsh Boechera selbyi var. inyoensis (Rollins) N. H. Holmgren
Treatment appears in FNA Volume 7. Treatment on page 385. Mentioned on page 358, 361, 386, 407.

Perennials; short-lived; apomictic; caudex usually not woody. Stems usually 1 per caudex, arising from center of rosette near ground surface, (1–) 2.5–6.5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. Basal leaves: blade oblanceolate, 1–4 (–8) mm wide, margins entire, ciliate along petiole and proximally, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–10-rayed, 0.2–0.7 mm. Cauline leaves: (7–) 12–35, concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves densely pubescent. Racemes 10–65-flowered, usually unbranched. Fruiting pedicels divaricate-ascending to horizontal, straight or slightly curved downward, 5–15 mm, sparsely pubescent, trichomes appressed, branched. Flowers ascending at anthesis; sepals pubescent; petals lavender to purplish, 5–8 × 1.2–2 mm, glabrous; pollen spheroid. Fruits divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.7–6.5 cm × 1.5–2.2 mm; valves glabrous; ovules 74–134 per ovary; style 0.05–0.2 mm. Seeds usually sub-biseriate, rarely uniseriate, 1.7–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. 2n = 21.


Phenology: Flowering Apr–Jun.
Habitat: Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands
Elevation: 1400-2400 m

Discussion

Some plants here assigned to Boechera inyoensis were treated by R. C. Rollins (1993) and N. H. Holmgren (2005b) as Arabis (or B.) beckwithii. Closer examination of the type specimens of A. beckwithii reveals that they are conspecific with B. puberula, a sexual diploid widespread in the Great Basin of western North America. Boechera inyoensis is an apomictic triploid that clearly contains at least one genome derived from B. shockleyi (M. D. Windham and I. A. Al-Shehbaz 2007b); the other parent has not been determined.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera inyoensis"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Rollins) Al-Shehbaz +
not +  and rosulate +
Arabis inyoensis +
appendaged +  and unappendaged +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Calif. +, Nev. +  and Utah. +
3.7 cm37 mm <br />0.037 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1400-2400 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
straight +  and curved +
divaricate-ascending +  and widely pendent +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
lavender +  and purplish +
rudimentary +
oblanceolate +  and spatulate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Apr–Jun. +
trinucleate +  and 3(-11)-colpate +
spheroid +
unbranched;10-65-flowered +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
sub-biseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
orbicular +, oblong +  and flattened +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
Arabis holboellii var. derensis +  and Boechera selbyi var. inyoensis +
Boechera inyoensis +
Boechera +
species +
branched +  and short-stalked +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
curved +  and straight +
perennial +  and short-lived +
downward +, divaricate-ascending +  and horizontal +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.1mm;0.2mm +