Boechera horizontalis

(Greene) Windham & Al-Shehbaz

Harvard Pap. Bot. 11: 266. 2007.

Basionym: Arabis horizontalis Greene Leafl. Bot. Observ. Crit. 2: 74. 1910
Synonyms: Arabis suffrutescens var. horizontalis (Greene) Rollins
Treatment appears in FNA Volume 7. Treatment on page 384. Mentioned on page 360, 361.

Perennials; long-lived; (cespitose); apomictic; caudex woody. Stems usually 1 per caudex branch, arising from center of rosette, near ground surface or somewhat elevated on woody base, or 1–3.5 dm, sparsely pubescent proximally, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm, glabrous distally. Basal leaves: blade narrowly oblanceolate, 1–5 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm. Cauline leaves: 3–13, often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. Racemes 5–32-flowered, usually unbranched. Fruiting pedicels horizontal to descending, straight or slightly curved downward, 4–11 mm, glabrous. Flowers divaricate-ascending at anthesis; sepals pubescent; petals lavender to purple, 5–6 × 1.5–2 mm, glabrous; pollen spheroid. Fruits horizontal or descending, not appressed to rachis, secund, straight, edges slightly undulate (not parallel), 2–4 cm × 2–3 mm; valves glabrous; ovules 40–54 per ovary; style 0.2–0.5 mm. Seeds uniseriate, 2–2.5 × 1.7–2 mm; wing continuous, 0.5–1 mm wide.


Phenology: Flowering Jul–Aug.
Habitat: Dry pumice slopes

Discussion

Morphological evidence suggests that Boechera horizontalis is an apomictic species that arose through hybridization between B. lemmonii and B. suffrutescens (M. D. Windham and I. A. Al-Shehbaz 2007). Boechera horizontalis is known only from the vicinity of Crater Lake in south-central Oregon.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera horizontalis"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Greene) Windham & Al-Shehbaz +
not +  and rosulate +
Arabis horizontalis +
appendaged +  and unappendaged +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
undulate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
divaricate-ascending +
straight +
descending +  and horizontal +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Dry pumice slopes +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
lavender +  and purple +
rudimentary +
oblanceolate +  and spatulate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Jul–Aug. +
trinucleate +  and 3(-11)-colpate +
spheroid +
Harvard Pap. Bot. +
unbranched;5-32-flowered +
curved +  and straight +
downward +, horizontal +  and descending +
slender +
0.4 cm4 mm <br />0.004 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
mucilaginous +
margined +, not winged +  and winged +
biseriate +, sub-biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
orbicular +, oblong +  and flattened +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
glabrous +  and pubescent +
Arabis suffrutescens var. horizontalis +
Boechera horizontalis +
Boechera +
species +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and long-lived +
hispid +, hirsute +, pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.5mm;1mm +